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In Tochigi Prefecture, none of 17 adult overwintering H. longicornis showed theilerial bodies in the salivary glands. However 19 of 186 nymphs did show infection with increasing numbers and amount of protozoan masses from October to February. A splenectomized calf exposed to 40 overwintering nymphs ...

Fujisaki, K.; Ito, Y.; Kamio, T.; Kitaoka, S.
Annals of Tropical Medicine and Parasitology, 1985, 79, 5, pp 519-524