Ambloplites rupestris (rock bass)

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Title

Caption

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Adult

Ambloplites rupestris (rock bass); adult. Captive fish in the Aquarium du Québec, Canada. April, 2012.

©Simon Pierre Barrette (cephas)-2012 via wikipedia - CC BY-SA 3.0

Identity

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Preferred Scientific Name

Ambloplites rupestris Rafinesque

Preferred Common Name

rock bass

Other Scientific Names

Ambloplites rupestris Jordan and Gilbert
Bodianus rupestris Rafinesque
Centrarchus aeneus Richardson
Cichla aenea Lesueur

International Common Names

English: goggle-eye; red eye; rock perch

Local Common Names

Canada: northern rock bass; redeye bass
Denmark: stenaborre; stenbars
Finland: kiviahven
France: crapet de roche
Germany: gemeiner felsenbarsch; germeiner sonnenbarsch; sonnenfisch; steinbarsch
Poland: bass czerwonooki
Portugal: perca-da-rocha
Sweden: stenabborre

Summary of Invasiveness

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A. rupestris, commonly known as rock bass, is a member of the sunfish family. A. rupestris has been introduced throughout the USA and into countries outside of North America, typically as a result of the sport-fishing industry. The limited research available shows that A. rupestris feeds on aquatic plants, insects and small fish. As a result it competes with other native species for food resources. In addition to this it is possible for A. rupestris to hybridize with roanoke (A. cavifrons) and shadow bass (A. ariommus). Roanoke bass is a species of special concern in North Carolina and Virginia (Jenkins and Burkhead, 1994).

Taxonomic Tree

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Domain: Eukaryota
Kingdom: Metazoa
Phylum: Chordata
Subphylum: Vertebrata
Class: Actinopterygii
Order: Perciformes
Suborder: Percoidei
Family: Centrarchidae
Genus: Ambloplites
Species: Ambloplites rupestris

Notes on Taxonomy and Nomenclature

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A.rupestris was first described by Rafinesque in 1817 and is a member of the sunfish family.

The definition of Ambloplites - meaning “blunt shield” and rupestris – preference for being amongst rocks (Schnell, 2014).

Description

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A. rupestris is a member of the sunfish family, and is characterized with a deep body, having its greatest depth at the origin of the dorsal fin being 31.8-36.7% of its total length (Scott and Crossman, 1973). Similar to other sunfish this fish is laterally compressed with the dorsal fin consisting of two confluent parts, an anterior part consisting of eleven sharp spines, with a posterior part of soft rays. The anal fin has six spines, with this number allowing the identification of rock bass from other members of the family. The iris of the eye is usually bright red to orange. A. rupestris can vary in colour and has been described as various shades of olive green with brassy or coppery reflections with a little black spot on the tip of each scale giving the appearance of a series of stripes the full length of the body.

Juvenile A. rupestris are pale or yellow-green irregularly barred and blotched with black.

Distribution

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The native range of A. rupestris is restricted to the fresh waters of east-central North America and can be found from southern Ontario, the Great Lakes and St. Lawrence River to Quebec and Lake Champlain; found west of the Appalachians south to the Tennessee River drainage and in tributaries of the middle and upper Mississippi River valley. A. rupestris has however been introduced throughout the USA into areas including Atlantic slope drainages (Scott and Crossman, 1973; Cashner, 1980; Cashner and Jenkins, 1982).

There is some disagreement with regards to the status of A. rupestris in Arkansas, Nebraska and Oklahoma with reports of this species being both introduced and native to these areas (Jones 1963; Scott and Crossman, 1973; Lee et al., 1980; Cross et al., 1986; USGS NAS, 2015).

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 17 Dec 2021

Continent/Country/Region	Distribution	Origin	First Reported	Reference	Notes
Europe
Czechia	Absent, Formerly present		1892	Seebens et al. (2017)
France	Present, Localized	Introduced		Keith and Allardi (2001) Seebens et al. (2017)	Established, reproducing naturally in Loire
Germany	Present	Introduced		Seebens et al. (2017)	First reported: 1880 - 1889
United Kingdom	Present, Localized	Introduced		Welcomme (1988) Seebens et al. (2017)	Released into a lake in Oxford, likely extirpated
North America
Canada	Present			CABI (Undated a)	Present based on regional distribution.
-Manitoba	Present, Widespread	Native		Scott and Crossman (1973)
-Ontario	Present, Widespread	Native		Scott and Crossman (1973)
-Quebec	Present, Widespread	Native		Scott and Crossman (1973)
-Saskatchewan	Present, Widespread	Native		Scott and Crossman (1973)
Mexico	Present	Introduced		Welcomme (1988)
United States	Present			CABI (Undated a)	Present based on regional distribution.
-Alabama	Present	Native		Page and Burr (1991)
-Arizona	Present, Localized	Introduced		Miller and Lowe (1967) USGS NAS (2014)
-Arkansas	Present			Scott and Crossman (1973) Lee et al. (1980) Cross et al. (1986) Robison and Buchanan (1988) USGS NAS (2014)	Recorded as both introduced and native
-California	Present, Localized	Introduced		Smith (1896) Dill and Cordone (1997) USGS NAS (2014)
-Colorado	Present	Introduced		Scott and Crossman (1973) USGS NAS (2014)
-Delaware	Present, Localized	Introduced		Raasch and Altemus (1991) USGS NAS (2014)
-District of Columbia	Present, Localized	Introduced		Tilmant (1999) USGS NAS (2014)
-Florida	Present	Native		Scott and Crossman (1973)
-Georgia	Present	Native		Scott and Crossman (1973)
-Idaho	Present, Localized	Introduced		Linder (1963)
-Indiana	Present	Introduced		Sweeney (1902)
-Kansas	Present	Introduced		Cross (1967) Scott and Crossman (1973) Cross and Collins (1995)
-Maryland	Present, Localized	Introduced		Lee et al. (1980) Tilmant (1999) Rhode et al. (2009) CABI (Undated)
-Massachusetts	Present, Localized	Introduced		Hartel (1992)
-Missouri	Present, Localized	Native		Page and Burr (1991)
-Montana	Present, Localized	Introduced		Cross (1967)
-Nebraska	Present			Jones (1963) Scott and Crossman (1973)	Recorded as both introduced and native
-New Hampshire	Present, Localized	Introduced		Scarola (1973)
-New Jersey	Present, Localized	Introduced		Nelson (1890)
-New Mexico	Present, Localized	Introduced		Sublette et al. (1990)
-North Carolina	Present, Localized	Introduced		Menhinick (1991)
-North Dakota	Present	Native		Scott and Crossman (1973)
-Oklahoma	Present			Scott and Crossman (1973) Lee et al. (1980)	Recorded as both introduced and native
-Pennsylvania	Present, Localized	Introduced		Lee et al. (1980) Tilmant (1999)
-South Carolina	Present, Localized	Introduced		Rhode et al. (2009)
-South Dakota	Present, Localized	Introduced		Bailey and Allum (1962)
-Texas	Present, Localized	Introduced		Hubbs et al. (1991) Rasmussen (1998)
-Utah	Present, Localized	Introduced		Sigler and Sigler (1996)
-Virginia	Present, Localized	Introduced		Lee et al. (1980) CABI (Undated)
-Washington	Present, Localized	Introduced		CABI (Undated)	Original citation: Beecher Fernau (1982)
-West Virginia	Present, Localized	Introduced		Stauffer et al. (1995)
-Wyoming	Present	Introduced		Scott and Crossman (1973)

History of Introduction and Spread

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A. rupestris has been intentionally stocked within the USA for over a century. From 1889 to 1936, the United States Fish Commission introduced rock bass to Atlantic drainages and some western states. Research suggests that this species was intentionally stocked as a sport-fishing target with its expansion through natural dispersal. Outside of the USA and Canada, this species was introduced to Mexico as an aquaculture species (Page and Burr, 1991).

Similar to its introduction in the USA, this species was introduced into a lake in the UK; Linkside Lake, Oxford, during the period of 1925-1949 with a possible second attempt at introduction in the 1990s (Welcomme, 1988). It is believed that this species is still present (Wheeler and Maitland, 1973), although attempts to survey have been unsuccessful.

In France, A. rupestris has been the subject of several introduction attempts from the end of the nineteenth century up until the mid-twentieth century in the Saone and Loire regions. Today, only one population of A. rupestris appears to be present in Loire, with its distribution apparently on the increase (Achigan, 2015).

A. rupestris was introduced into Berneuchen, Germany in 1887 from the USA (von dem Borne, 1890; von Wengen, 1892). A total of 20 fish were imported and by 1889 12 of these were still alive and had successfully spawned. Nevertheless A. rupestris is no longer present in Germany (de Groot, 1985).

In 1892 A. rupestris was also introduced into the Czech Republic as an experiment, which was ultimately unsuccessful (Lusk et al., 2010).

Introductions

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Introduced to	Introduced from	Year	Reason	Introduced by	Established in wild through		References	Notes
Introduced to	Introduced from	Year	Reason	Introduced by	Natural reproduction	Continuous restocking	References	Notes
France	North America	1904-1910			Yes	No	Keith and Allardi (2001)	Research
Germany		1880-1889			No	No	NOBANIS (2013)
Mexico	USA		Aquaculture (pathway cause)		Yes	No	Welcomme (1988)
UK	USA	1930	Stocking (pathway cause)		Yes	No	Welcomme (1988)	Potentially extinct

Risk of Introduction

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The spread of A. rupestris into lakes, reservoirs and streams in new areas is often a result of intentional introduction of this fish as a sport fish, intentional aquarium releases and for aquaculture.

Habitat

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A. rupestris generally inhabits rocky, vegetated areas in shallow water such as lakes, brushy stream margins, pools of creeks and small to medium rivers (Page and Burr, 1991). They usually inhabit freshwater with a high cover of vegetation to provide protection from predators.

In Europe, A. rupestris has a tendency to avoid swift waters and occurs in a wide variety of slow-flowing to stagnant waters such as large rivers, lakes, ponds, canals and backwaters (Kottelat and Freyhof, 2007).

Habitat List

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Category	Habitat	Presence	Status
Freshwater
Freshwater	Irrigation channels	Secondary/tolerated habitat	Natural
Freshwater	Lakes	Principal habitat	Natural
Freshwater	Reservoirs	Principal habitat	Natural
Freshwater	Rivers / streams	Principal habitat	Natural
Freshwater	Ponds	Secondary/tolerated habitat	Natural

Biology and Ecology

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Genetics

A.rupestris has a diploid (2n) chromosome number of 48 and a haploid/gametic (n) of 24 (Arkhipchuk, 1999).

Reproductive Biology

A. rupestris are polygyandrous; both males and females have several mates during the breeding season (Schnell, 2014). In its native range in North America, A. rupestris spawn in late spring and early summer when water temperatures reach 15.6-21.1°C (Scott and Crossman, 1973). Males dig shallow nests, where spawning can take place with several females over a period of time. Depending on the size of the female, 500-5,000 eggs may be laid in one nest (Schnell, 2014). After fertilization the male defends the eggs from predators and cares for the young; a total of about 14 days. Both male and females reach maturity at two to three years of age (Schnell, 2014).

Outside of its native range, males fan and defend the nests, which are shallow depressions in sand or gravel bottom along shallow shores.

Longevity

A. rupestris has an average lifespan of about five to eight years in the wild. It has been reported that the maximum lifespan of a rock bass in captivity was 18 years (Altman and Dittmer, 1962).

Nutrition

A. rupestris feeds on aquatic plants, insects and small fish, including young of their own species (Page and Burr, 1991).

Associations

A. rupestris have been known to carry a number of parasites, which include: Protoza, Trematoda, Cestoda, Nematoda, Acanthocephala, leeches and Crustacea (Hoffman, 1967). One parasite regularly identified on this species is “black-spot” (Neascus species) (Scott and Crossman, 1973).

Environmental Requirements

A. rupestris can survive in areas with a broad temperature range <10-29°C) (Schnell, 2014). It has been suggested that it has a preferred range of 21-26°C.

Natural Food Sources

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Food Source	Food Source Datasheet	Life Stage
Amphipoda		All Stages
Anisoptera		All Stages
	Anisoptera
Chironomids		All Stages
Cladocera		All Stages
Copepoda		All Stages
Crayfish		All Stages
Ephemeroptera		All Stages
Hydracarina		All Stages
Isopoda		All Stages
Odonata		All Stages
Small fish species		Aquatic\|Broodstock; Aquatic\|Fry; Aquatic\|Larval
Trichoptera		All Stages
yellow perch		Aquatic\|Larval

Climate

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Climate	Status	Description
D - Continental/Microthermal climate	Preferred	Continental/Microthermal climate (Average temp. of coldest month < 0°C, mean warmest month > 10°C)
Df - Continental climate, wet all year	Preferred	Continental climate, wet all year (Warm average temp. > 10°C, coldest month < 0°C, wet all year)
Ds - Continental climate with dry summer	Preferred	Continental climate with dry summer (Warm average temp. > 10°C, coldest month < 0°C, dry summers)
Dw - Continental climate with dry winter	Preferred	Continental climate with dry winter (Warm average temp. > 10°C, coldest month < 0°C, dry winters)

Water Tolerances

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Parameter	Minimum Value	Maximum Value	Typical Value	Status	Life Stage	Notes
Dissolved oxygen (mg/l)			> 6	Optimum

Notes on Natural Enemies

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There are several predators in the native range which prey on young A. rupestris. These include largemouth bass (Micropterus salmoides), northern pike (Esox Lucius), and muskellunge (Esox masquinongy).

Means of Movement and Dispersal

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Accidental Introduction

Accidental introductions of A. rupestris have been reported through bait bucket releases.

Intentional Introduction

In the past, A. rupestris has been stocked as an aquarium species as well as an angling target for sport fishing. Intentional introduction is the main cause of introduction of A. rupestris into new areas, particularly those over longer distances.

Pathway Causes

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Cause	Long Distance	Local
Aquaculture	Yes	Yes
Fisheries	Yes	Yes
Hunting, angling, sport or racing	Yes	Yes
Intentional release	Yes	Yes

Pathway Vectors

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Vector	Notes	Long Distance	Local	References
Pets and aquarium species		Yes	Yes
Water		Yes	Yes

Impact Summary

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Category	Impact
Economic/livelihood	Positive
Environment (generally)	None

Economic Impact

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There is limited economic impact associated with A. rupestris. This species is not highly sought after as a sports-fish nor a commercial species, but has some value as an aquarium species.

Environmental Impact

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Impact on Biodiversity

A. rupestris competes with native species and is reported to have severely affected populations of roanoke bass (Ambloplites cavifrons) through hybridization and competition (Lee et al., 1980; Jenkins and Burkhead, 1994). Roanoke bass are a species of special concern in North Carolina and Virginia (Jenkins and Burkhead, 1994). It is also know to hybridize with shadow bass (A. ariommus).

A. rupestris have also been shown to reduce the abundance and diversity of native prey species in several Pacific Northwest rivers (Hughes and Herlihy, 2012).

A. rupestris have been shown to reduce the abundance and diversity of native prey species in several Pacific Northwest rivers (Hughes and Herlihy, 2012). Jenkins and Burkhead (1994) speculated that introduced A. rupestris have contributed to the demise of an isolated population of trout-perch (Percopsis omiscomaycus) in the Potomac River in Virginia and Maryland. Additionally, research shows a decline in Smallmouth bass (Micropterus dolomieu) populations where A. rupestris has been introduced as a result of competition for food resources, specifically minnows and crayfish. It has also been found to compete with walleye (Sander vitreus) for food resources and may also predate on young walleye. Both smallmouth bass and walleye are highly sought after fish as recreational targets.

Threatened Species

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Threatened Species	Conservation Status	Where Threatened	Mechanism	References	Notes
Etheostoma nianguae (Niangua darter)	VU (IUCN red list: Vulnerable); USA ESA listing as threatened species	Missouri	Competition - monopolizing resources	US Fish and Wildlife Service (1989)
Gila nigrescens (chihuahua chub)	VU (IUCN red list: Vulnerable); USA ESA listing as threatened species	New Mexico	Competition - monopolizing resources	US Fish and Wildlife Service (2010)

Risk and Impact Factors

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Invasiveness

Invasive in its native range
Proved invasive outside its native range
Has a broad native range
Abundant in its native range
Tolerant of shade
Capable of securing and ingesting a wide range of food
Highly mobile locally

Impact outcomes

Altered trophic level
Conflict
Negatively impacts aquaculture/fisheries
Reduced native biodiversity
Threat to/ loss of endangered species
Threat to/ loss of native species

Impact mechanisms

Competition - monopolizing resources
Competition - smothering
Pest and disease transmission
Hybridization
Predation

Likelihood of entry/control

Highly likely to be transported internationally deliberately
Highly likely to be transported internationally illegally

Uses

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Economic Value

In some areas, A. rupestris is considered a valuable sport fish, with bass fishing being a multi-billion dollar industry in the USA. Fishing supports approximately 828,000 jobs in the USA; many of these jobs involve fishing for species such as rock bass (Schramm et al., 1991).

Social Benefit

A. rupestris is an important fish species for recreational fishery.

Uses List

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General

Pet/aquarium trade
Sport (hunting, shooting, fishing, racing)

Human food and beverage

Meat/fat/offal/blood/bone (whole, cut, fresh, frozen, canned, cured, processed or smoked)

Detection and Inspection

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A. rupestris can be easily identified from other species of sunfish by the presence of six spines on the anal fin and a bright red to orange eye.

Similarities to Other Species/Conditions

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A. rupestris, otherwise known as rock bass, aren’t actually bass and are instead members of the sunfish family. Confusion between other members of the sunfish family is often with similarities between the bluegill (Lepomis macrochirus) and warmouth (Lepomis gulosus); however, A. rupestris can be readily distinguished from these species by the six spines in the anal fin (other typical sunfishes have only three anal fin spines).

Prevention and Control

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Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.

Prevention

Public Awareness

Public awareness is important in preventing establishment of new populations and further introductions of A. rupestris.

Eradication

Annual rock bass fishing tournaments are held in the USA to aid in eradication of this species.

Control

Physical/Mechanical Control

Electrofishing and seine/gill netting has been used to catch A. rupestris in its native ranges, however there has been no report to use these methods to physically or mechanically control this species.

Chemical Control

The only effective method eradicating A. rupestris is by the application of rotenone, a piscicide. This chemical however, is also toxic to non-target species.

Monitoring and Surveillance

Both radio and acoustic telemetry can be used to detect A. rupestris.

Gaps in Knowledge/Research Needs

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There is little information on the potential and current impacts of A. rupestris in its introduced range.

References

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Achigan, 2015. (Peche sportive). [English title not available]. France. http://www.achigan.net/en/

Altman PL, Dittmer DS, 1962. Growth, including reproduction and morphological development. Federation of American Societies for Experimental Biology, Washington, xii + 608 pp.

Arkhipchuk V, 1999. Chromosome database. Database of Dr. Victor Arkhipchuk.

Bailey RM, Allum MO, 1962. Fishes of South Dakota. Michigan, USA: Miscellaneous Publications, Museum of Zoology, University of Michigan, 119.

Beecher HA, Fernau RF, 1982. Fishes of Oxbow Lakes of Washington. Northwest Science, 57(2):125-131.

Borne M, dem von, 1890. Der Amerikanische Steinbarsch (Rock-Bass) in Deutschland ([English title not available]). Neudamm, Poland: Verlag J. Neumann.

Cashner RC, 1980. Ambloplites rupestris (Rafinesque), rock bass. In: Atlas of North American Fishes [ed. by Lee, D. S. \Gilbert, C. \Hocutt, C. \Jenkins, R. \McAllister, D. E. \Stauffer, J. R.]. Raleigh, North Carolina, USA: NC State Museum of Natural History, 581.

Cashner RC, Jenkins RE, 1982. Systematics of the Roanoke bass, Ambloplites cavifrons. Copeia, 3:581- 594.

Cross FB, 1967. Handbook of fishes of Kansas. Kansas, USA: Museum of Natural History, University of Kansas, 375 pp. [Miscellaneous publication, no. 45.]

Cross FB, Collins JT, 1995. Fish in Kansas, second edition. Lawrence Kansas, USA: Univeristy of Kansas Museum of Natural History Public Education.

Cross FB, Mayden RL, Stewart JD, 1986. Fishes in the western Mississippi basin (Missouri, Arkansas, and Red Rivers). In: The Zoogeography of North American Freshwater Fishes [ed. by Hocutt CH, Wiley EO, ] New York, USA: John Wiley and Sons, 363-412.

Dayton P, Thrush S, Agardy T, Hofman R, 1995. Environmental effects of marine fishing. Aquatic Conservation: Marine and Freshwater Ecosystems, 5/3:205-232.

Dill WA, Cordone AJ, 1997. History and status of introduced fishes in California. Fish Bulletin, California Department of Fish and Game, 178:1-414.

Groot SJ de, 1985. Introductions of non-indigenous fish species for release and culture in the Netherlands. Aquaculture, 46:237-257.

Hartel K, 1992. Non-native fishes known from Massachusetts freshwaters. Occasional Reports of the MCZ Fish Department, 1992(2). 1-9.

Hoffmann GL, 1967. Parasites of North American Freshwater Fishes. Berkeley, California, USA; University of California Press, 486 pp.

Hubbs C, Edwards RJ, Garrett GP, 1991. An annotated checklist of freshwater fishes of Texas, with key to identification of species, 43(4):1-56.

Hughes RM, Herlihy AT, 2012. Patterns in catch per unit effort of native prey fish and alien piscivorous fish in 7 Pacific Northwest USA rivers. Fisheries (Bethesda), 37(5):201-211.

Jenkins RE, Burkhead NM, 1994. Freshwater fishes of Virginia. Bethesda, MD, USA: American Fisheries Society.

Jones DJ, 1963. A history of Nebraska's fishery resources. Nebraska, USA: Nebraska Game, Forestation and Parks Commission.

Keith P, Allardi J, 2001. Atlas des poissons d'eau douce de France (Atlas of the freshwater fishes of France), 387 pp. [Patrimoines naturels, No. 47]

Kottelat M, Freyhof J, 2007. Handbook of European Freshwater Fishes. Cornol, Switzerland: Publications Kottelat, 646 pp.

Lee DS, Gilbert CR, Hocutt CH, Jenkins RE, McAllister DE, Stauffer JRJr, 1980. Atlas of North American freshwater fishes. Raleigh, NC, RC: North Carolina State Museum of Natural History,.

Linder AD, 1963. Idaho's alien fishes. Journal of the Idaho Museum of Natural History, 6(2):12-15.

Lusk S, Lusková V, Hanel L, 2010. Alien fish species in the Czech Republic and their impact on the native fish fauna. Folia Zoologica, 59(1):57-72.

Menhinick EF, 1991. The freshwater fishes of North Carolina. Raleigh, North Carolina, USA: North Carolina Wildlife Resources Commission, 227.

Miller RR, Lowe CH, 1967. Fishes of Arizona. In: The Vertebrates of Arizona [ed. by Lowe, C. H.]. Tucson, Arizona, USA: University of Arizona Press, 133-151.

Missouri Department of Conservation, 2008. Fish of Missouri. Missouri, USA: Missouri Department of Conservation. http://mdc.mo.gov/

Nelson J, 1890. Discriptive catalogue of the vertebrates of New Jersey. Geological Survery of New Jersey 1890. 489-824.

NOBANIS, 2013. North European and Baltic Network on Invasive Alien Species. http://www.nobanis.org/

Page LM, Burr BM, 1991. A field guide to freshwater fishes of North America north of Mexico. Boston, USA: Houghton Mifflin Company, 432 pp.

Raasch MS, Altemus VL, 1991. Delaware's freshwater and brackish water fishes - a popular account. Delaware, USA: Delaware State College, Center for the Study of Del-Mar-Va Habitats, 166 pp.

Rasmussen JL, 1998. Aquatic nuisance species of the Mississippi river basin. In: 60th Midwest Fish and Wildlife Conference, Aquatic Nuisance Species Symposium, Dec. 7, 1998, Cincinnati, OH. unpaginated.

Rhode FC, Arndt RG, Foltz JW, Quattro JM, 2009. Freshwater fishes of South Carolina. Columbia, South Carolina, USA: University of South Carolina Press, 430 pp.

Robinson HW, Buchanan TM, 1988. Fishes of Arkansas. Arkansas, USA: University of Arkansas Press, 536.

Scarola JF, 1973. Freshwater Fishes of New Hampshire. New Hampshire, USA: New Hampshire Fish and Game Department, Division of Inland and Marine Fisheries, 131.

Schnell B, 2014. Ambloplites rupestris, Goggle eye. Michigan, USA: University of Michigan. http://animaldiversity.org/accounts/Ambloplites_rupestris/

Schramm Jr H, Armstrong M, Funicelli N, Green G, Lee D, Manns Jr R, Taubert B, Waters S, 1991. The status of competitive sport fishing in North America. Fisheries, 16/3:4-12.

Scott WB, Crossman EJ, 1973. Freshwater fishes of Canada. Bull. Fish. Res. Board Can. 184:1-966.

Sigler WF, Sigler JW, 1996. Fishes of Utah: a natural history. Salt Lake City, Utah, USA: University of Utah Press.

Smith HM, 1896. A review of the history and results of the attempts to acclimatize fish and other water animals in the Pacific states. Bulletin of the US Fish Commission, 15:379-472.

Stauffer JR, Boltz JM, White LR, 1995. The fishes of West Virginia. The Proceedings of the Academy of Natural Sciences of Philadelphia, 146:1-389.

Sublette JE, Hatch MD, Sublette M, 1990. The fishes of New Mexico. Albuquerque, New Mexico: University New Mexico Press, 393 pp.

Sweeney ZT, 1902. Biennial report of the commissioner of fisheries and game for Indiana. Indianapolis, Indiana, USA: WM. B. Burford.

Tilmant JT, 1999. Management of nonindigenous aquatic fish in the U.S. National Park System. USA: National Park Service, 50 pp.

US Fish and Wildlife Service, 1989. In: A Recovery Plan for the Niangua Darter (Etheostoma nianguea). US Fish and Wildlife Service, 44 pp.. http://www.fws.gov/ecos/ajax/docs/recovery_plan/890717.pdf

US Fish and Wildlife Service, 2010. In: Chihuahua Chub (Gila nigrescens). 5-Year Review: Summary and Evaluation. US Fish and Wildlife Service, 23 pp.. http://ecos.fws.gov/docs/five_year_review/doc4325.pdf

USGS NAS, 2014. Nonindigenous aquatic species database. Gainesville, Flordia, USA: USGS. http://nas.er.usgs.gov/

Welcomme RL, 1988. International introductions of inland aquatic species. FAO Fisheries Technical Paper No. 294. Rome, Italy: FAO, 318 pp.

Wengen FR von, 1892. Neue Amerikanische Fische in Deutschland. Circulare des Deutschen Fischerei-Vereins vom 1892. Berlin, Germany: W. Moseser, 236-237.

Wheeler AC, Maitland PS, 1973. The scarcer freshwater fishes of the British Isles I. Introduced species. Journal of Fish Biology, 5:49-68.

Distribution References

Bailey RM, Allum MO, 1962. Fishes of South Dakota., Michigan, USA: Miscellaneous Publications, Museum of Zoology, University of Michigan. 119.

CABI, Undated. Compendium record. Wallingford, UK: CABI

CABI, Undated a. CABI Compendium: Status inferred from regional distribution. Wallingford, UK: CABI

CABI, Undated b. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI

Cross F B, 1967. Handbook of fishes of Kansas. Kansas, USA: Museum of Natural History, University of Kansas. 375 pp.

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12/02/15 Original text by:

Michael Godard, consultant, Canada

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Ambloplites rupestris (rock bass)

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