Solenopsis geminata
(tropical fire ant)

Pictures

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Identity

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Preferred Scientific Name

• Solenopsis geminata (Fabricius)

Preferred Common Name

• tropical fire ant

Other Scientific Names

• Atta geminata
• Solenopsis geminata rufa Jerd.

International Common Names

• English: brown ant; fire ant; fire ant, tropical; red ant; stinging ant
• Spanish: hormiga brava; hormiga brava roja; hormiga caribe
• French: fourmie

Local Common Names

• Netherlands: Tabaks-mier

EPPO code

• SOLEGE (Solenopsis geminata)
• SOLEGR (Solenopsis geminata rufa)

Summary of Invasiveness

Top of page Relative to other invasive fire ant species such as Solenopsis invicta, the tropical fire ant, S. geminata, has a greater world-wide distribution. Where introduced, it has often become one of the dominant pest ant species, affecting fauna and flora (Hoffmann et al., 1999; Geetha Viswanathan and Ajay Narendra, 2000).

Taxonomic Tree

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• Domain: Eukaryota
•     Kingdom: Metazoa
•         Phylum: Arthropoda
•             Subphylum: Uniramia
•                 Class: Insecta
•                     Order: Hymenoptera
•                         Family: Formicidae
•                             Genus: Solenopsis
•                                 Species: Solenopsis geminata

Notes on Taxonomy and Nomenclature

Top of page Taber (2000) provided the historical background of the taxonomic status of the tropical fire ant, S. geminata. The fire ants comprise a group of 18 to 20 species that are native to the New World and within the genus Solenopsis. This genus contains approximately 200 species including 'thief ants', most of which are incapable of stinging humans, and nest close to other ant nests and rob them of stored food and brood. Trager (1991) recently revised this group of ants. Several colour forms have been recognized (Hölldobler and Wilson, 1990). Barrion et al. (1979) refer to the subspecies Solenopsis geminata rufa.

Description

Top of page Hung et al. (1977) provided a key for the identification of fire ants in Texas, USA. S. geminata worker ants have two nodes between the thorax and abdomen, distinct compound eyes, ten segments on the antennae with two segments forming a club, no spines on the propodeum and are over 2 mm long. Major worker ants have a deep groove on the vertex of the head, mandibles without teeth and are entirely black, with a short antennal scape reaching half-way to the vertex (Vinson et al., 2003).

Distribution

Top of page This species is native to the New World, including the Caribbean islands such as Puerto Rico. It has spread world-wide, in some cases to environments where the conditions are unfavourable for this species to survive outdoors such as Winnipeg, Canada (Ayre, 1977).

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 23 Apr 2020

History of Introduction and Spread

Top of page Taber (2000) provides a world-wide distribution map for S. geminata. This species has been widely spread through commerce to new locations in North and South America, Africa, Australia, Asia and the islands of Oceania, where it can become a dominant species (George and Narendran, 1987; Porter et al., 1997). This is a predominantly tropical species.

In parts of southern USA, the historical distribution of S. geminata has been reduced in areas invaded by high population levels of imported fire ants (Solenopsis invicta and Solenopsis richteri) (Vinson et al., 2003). Thus, historical records may no longer express the current distribution.

Risk of Introduction

Top of page The phytosanitary risk is high for areas that do not already have this species and are located in warmer climates.

Habitat

Top of page S. geminata generally nests in the soil of suitable environments.

Hosts/Species Affected

Top of page S. geminata is omnivorous, but among the fire ants is known more as a seed-feeding species. The morphologically distinctive major worker ants are specialized in collecting and feeding on the seeds of many plant species, including weeds (Wilson 1978; Carroll and Risch, 1984; Tennant and Porter 1991; Sangal and Neena Bakshi, 1994). Although this ant is reported to occur in a number of crops (avocado, banana, cabbage, citrus, cocoa, corn, cotton, coffee, cucumber, eggplant, mango, okra, papaya, pastureland, pineapple, sorghum, soybeans, strawberries, sugarcane, tobacco, tomato, rice and vegetables) and in cultivated land, it is seldom reported to be a direct pest of these crops.

Seed loss to crops such as cabbage, corn, sorghum and tomato can be significant. However, this species also girdles plants, imbibes sap, and bites branches, shoots, buds, flowers and fruits (Hill, 1975). The ants in nests that are built at the base of citrus tree trunks gnaw through the bark (Essig, 1926; Wolcott, 1933). Indirectly, S. geminata can negatively affect plant health by tending sucking insects (Homoptera), or in one case, prevent pollination (Carroll and Risch, 1983). However, because these ants sting, they can affect field workers that are sensitive to arthropod venom, and can affect equipment such as drip irrigation systems (Ota and Chang, 1981) and electrical equipment (Prins, 1985). In addition, they occur in urban areas (such as in turf and occasionally enter buildings), wildlife areas, forests and other sites suitable for the species.

Host Plants and Other Plants Affected

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Growth Stages

Top of page Flowering stage, Fruiting stage, Pre-emergence, Seedling stage, Vegetative growing stage

List of Symptoms/Signs

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Biology and Ecology

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The research of S. geminata is fairly extensive, particularly from locations where it is an introduced exotic species. In these locations, S. geminata often becomes one of the more dominant ant species, causing many problems similar to those caused by Solenopsis invicta (Porter et al., 1997).
The biology of S. geminata is similar to other closely related fire ant species. Wilson (1978) described caste behaviour. The head capsules of the major worker ants are distinctly larger and lobed relative to those of minor workers because of their specialized seed foraging and milling behaviour.

The colonies of S. geminata can either be monogyne, each containing a single queen ant, or polygyne, containing many reproductively-active queen ants (Adams et al., 1976; MacKay et al., 1990; Williams and Whelan, 1991).

Genetics

Ross et al. (1987) has evaluated the genetics of S. geminata and other fire ant species. This species has been known to hybridize with other closely related fire ant species; Morrison (2000) discusses the hybrid Solenopsis geminata x Solenopsis xyloni.

Physiology and Phenology

The pheromones known for S. geminata are discussed by Blum (1982) and Jaffe et al. (1985), and considerable work has been carried out to characterize the ant's venom (e.g. Battenfield et al., 1982).

Reproductive Biology

The winged reproductive male and female S. geminata leave the nests on mating or nuptial flights, late in the day or at dusk on days of or following rains during warmer periods of the year (Travis, 1941; Hung et al., 1977; Trager, 1991). Two forms of queen ants are recognized and have different mating strategies (McInnes and Tschinkel, 1995). Following mating, the female reproductive ants or queens drop their wings and dig a burrow to start a new colony. The queens begin laying eggs and can lay up to 1123 eggs per day (Travis, 1941). Development from egg to adult can take from 3 to 4 weeks and up to 2 months (Wheeler and Wheeler, 1955; Vargo, 1993). Egg hatch occurs in 14 to 17 days, larval development in 24 days to 6 weeks and the pupal stages last up to 19 days (Veeresh, 1990a). The worker ants include minor workers (2.6 mm larvae), major workers (5.2 mm larvae) and intermediate sized workers. Colony maturity occurs in 1 to 2 years (Wilson and Taylor, 1967). The colonies have been reported to contain up to 100,000 ants (Way et al., 1998), but other estimates are lower, generally ranging from 10,000 to 12,000 (Veeresh, 1990a, b).

S. geminata, like other ants, are social insects. This species generally builds nests in the soil in open, sunny areas, although they can also occur in the shade and inside structures. The nests are formed as craters or mounds in which the ants seek optimum temperatures for reproduction. At certain times of the year, the nests have multiple entrance holes (16 to 20 per nest) in mounds up to 2 ft wide and 12 inches tall (Van Pelt, 1958), although most mounds are not as tall or even crater-like. The subterranean foraging tunnels (up to 3 inches deep) extend from the nest in all directions and may be 100 ft long, with vertical tunnels extending up to 6 ft deep in the ground (Wheeler, 1910). The density of colonies in areas varies greatly and can range from 10 mounds per acre to 80 mounds per acre in native habitats (McInnes and Tschinkel, 1995), but in some areas the polygyne forms of the ant can reach up to 1000 mounds per acre (MacKay et al., 1990). In India, mound densities have been reported as 1500 mounds per acre (Veeresh, 1990a). The colonies frequently migrate to new locations (Lakshmikantha et al., 1996), migrating over 100 ft from the original nest locations.

Environmental Requirements

S. geminata often colonizes disturbed habitats (Risch, 1981; Perfecto, 1991). It is capable of colonizing most types of soils and media. It occurs in shaded orchards and woods (Essig, 1926; Wilson and Brown, 1958), as well as open areas. Habitat types vary greatly (Wheeler, 1910; Travis, 1941; Creighton, 1950; Van Pelt, 1958; Whitcomb et al., 1972; Buren et al., 1974; Moody et al., 1981; Moody and Franke, 1982; Carroll and Risch, 1984; Tschinkel, 1988; Verhaagh, 1991; Muniappan and Marutani, 1992; McInnes and Tshinkel, 1995; Way et al., 1998). This species prefers low to mid-elevations below 1500 ft (Forel, 1993; Perfecto, 1994), but has been reported to occur at 3000 ft (Smith, 1936). It prefers mild winter temperatures and high humidity (Snelling, 1963). Cokendolpher and Francke (1985) described the temperature preferences of S. geminata. The influence of microhabitats on distribution is discussed by Torres (1984). Foraging occurs in the temperature range of 77 to 90°F, with extreme temperature limits that prevent foraging below 36°F and above 122°F (George and Narendran, 1987).

Associations

Although S. geminata is reported to be associated with a wide variety of crop and animal commodities in the field, the impact of the ants is often not reported to be negative, nor is injury always caused to the agricultural commodity. In many cases, the foraging ants prey on pest arthropods such as ticks, caterpillars and beetle life stages. They are associated with sucking insects such as aphids, leafhoppers and other Homoptera, which they 'tend', to feed on the sugary honeydew secretions (Tennant and Porter, 1991).

Natural enemies

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Means of Movement and Dispersal

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S. geminata can be moved by the human transport of colonies or mated queen ants in soil or other suitable nesting materials, or naturally by mating flights or the movement of colonies floating in floodwater (Litsinger et al., 1986), similar to the ways in which Solenopsis invicta and Solenopsis richteri have been spread to new areas. The cracks and crevices of stems and bark could harbour mated queen ants of small colonies, particularly associated with soil or potting media. The ants or 'worked' soil would be visible to the naked eye. The movement by humans has led to infestations on many remote islands such as the Galapagos Islands (Williams and Whelan, 1991) and Guam (Schreiner and Nafus, 1988).

Agricultural Practices

S. geminata often colonizes disturbed habitats (Risch, 1981; Perfecto, 1991). The removal of rain forests or the cultivation of fields causes rapid colonization by this species, making it a dominant ant in many agricultural systems.

Pathogen Transmission

S. geminata and other ant species may also be associated with the transmission of some plant diseases through the injuries made by the ant (Suarez-Sotolongo, 1990).
 

Pathway Vectors

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Plant Trade

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Plant parts not known to carry the pest in trade/transport
Flowers/Inflorescences/Cones/Calyx
Fruits (inc. pods)
Leaves
Roots
Seedlings/Micropropagated plants
True seeds (inc. grain)
Wood

Wood Packaging

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Impact Summary

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Impact

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S. geminata is considered to be an economically important pest ant (Lakshmikantha et al., 2001) even in Florida, USA, this species' native habitat (Wojcik et al., 1976). Stand loss in agricultural crops can be significant where this species is abundant. Perfecto (1994) reported a loss of 90% of tomato seeds. The loss of corn and sorghum seeds and seedlings has also been reported (Carroll and Risch, 1984; Trabanino et al., 1989). However, the overall estimates of economic losses are unavailable. Economic benefits can also be provided by this species (Risch and Carroll, 1982); it has been documented to be a major predator of many other arthropod pests such as engorged ticks, Boophilus microplus (Barre et al., 1991) and fruit flies (Eskafi and Kolbe, 1990).

Environmental Impact

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The impact of S. geminata is both positive and negative. Nesting activities improve soil conditions (Carroll and Risch, 1983) and foraging can reduce weed seeds (Risch and Carroll, 1986) and pest arthropod populations.

Impact: Biodiversity

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S. geminata has often become one of the dominant pest ant species where it has been introduced, affecting the fauna and flora (Hoffmann et al., 1999; Geetha Viswanathan and Ajay Narendra, 2000). This species affects the seed of many plant species, moving and consuming them as discussed by Horvitz (1981), Nesom (1981) and others. In native areas (New World), subsequently invaded by imported fire ant species (Solenopsis invicta and Solenopsis richteri in the south-eastern USA), S. geminata populations (and its distribution) have been reduced (Vinson et al., 2003).

Threatened Species

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Social Impact

Top of page Because of the ability for S. geminata to bite and sting, this species is considered a pest of humans and animals. However, relative to the more aggressive imported fire ant species (Solenopsis invicta and Solenopsis richteri), it is less medically important.

Risk and Impact Factors

Top of page Impact mechanisms

• Predation

Uses

Top of page The presence of S. geminata can be beneficial. However, other than use in conservation biological control programmes, the ant has not been used for augmentative biological control.

Diagnosis

Top of page Suspect-ants should be collected in a vial of alcohol using forceps, cotton swabs or other suitable collecting methods that prevent the collector from being stung. Five to twenty-five specimens, particularly of the larger, major worker ants, should be collected and submitted to an identification specialist for species verification.

Detection and Inspection

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The methods used for detecting and inspecting other fire ant species, such as Solenopsis invicta, can be used for S. geminata. Both species build nests in the soil and create mounds with honey-comb-shaped galleries in which ants reside. These can be observed on the soil surface, particularly after rainy periods. Any sites where the soil appears to be 'worked' and formed into small granulate particles should be inspected for the presence of ants. This is particularly important during the inspection of articles of trade stacked on pallets or containing suitable media for ant colony nesting. However, baiting methods are more suitable for detecting ants in transported articles, or in locations or seasons when the ants do not construct visible mounds. Bait stations to monitor S. geminata, using attractive substances such as tuna fish, have been evaluated (Nestel and Dickschen, 1990; Islam et al., 2000).

The Pacific Invasive Ant Key (PIAKey) manual Pacific Invasive Ants Taxonomy Workshop Manual can both be used in identifying invasive ants in the Pacific region.

Similarities to Other Species/Conditions

Top of page S. geminata is similar to the red imported fire ant, Solenopsis invicta (see data sheet on Solenopsis invicta).

Prevention and Control

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Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.

The efforts to develop controls for this pest in southern USA were reported as early as the 1930s (Travis, 1938, 1939), suggesting that this species of fire ant was considered pestiferous even before the arrival of the imported species. However, relative to the control programmes developed for the red imported fire ant, Solenopsis invicta, fewer management programmes have been specifically developed for S. geminata. Certainly, because these are both fire ants (closely related and similar in biology) the control programmes developed for S. invicta should also be useful for control of S. geminata. In areas of the world where S. geminata has been imported, their populations may be greater and control justified (George and Narendran, 1987; Porter et al., 1997). In India, management options are presented by Veeresh (1990b).

Phytosanitary Measures

The inspection of articles of trade that can potentially harbour this species (e.g. nursery stock, sod, hay, field-working equipment or articles of trade associated with soil or other suitable nesting media) should be routine and can prevent the introduction and spread of this species.

Cultural Control and Sanitary Methods

Ota and Chang (1979, 1981) and Heinz and Carlson (1984) developed drip irrigation in Hawaiian sugarcane fields and assessed it as resistant to damage by foraging S. geminata.

Host-Plant Resistance

None has been reported.

Biological Control

Although numerous organisms have been identified as natural enemies of S. geminata, few of these have been used as a method of control.

Chemical Control

Neem extract (azadiractin) has recently been assessed as a treatment for S. geminata (Geetha Viswanathan et al., 2002).

Field Monitoring/Economic Threshold Levels

No economic threshold levels have been developed for S. geminata.

IPM Programmes

As a species of 'fire ant', pest management programmes developed for S. invicta can be applied to this species where it occurs and causes problems that justify suppression.

References

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Adams CT, Banks WA, Plumley JK, 1976. Polygyny in the tropical fire ant, Solenopsis geminata with notes on the imported fire ant, Solenopsis invicta. Florida Entomologist, 59(4):411-415

Aesch L von, Cherix D, 2005. Introduced ant species and mechanisms of competition on Floreana Island (Galápagos, Ecuador) (Hymenoptera: Formicidae). Sociobiology, 45(2):463-482. http://www.csuchico.edu/biol/Sociobiology/volume/sociobiologyv45n22005.html#16

Ahmed S, Khan MR, Ahmed M, Ghaffar A, 1989. Natural enemies of paddy leaf-roller (Cnaphalocrocis medinalis Gn.). Journal of Agricultural Research (Lahore), 27(1):71-76

APPPC, 1987. Insect pests of economic significance affecting major crops of the countries in Asia and the Pacific region. Technical Document No. 135. Bangkok, Thailand: Regional Office for Asia and the Pacific region (RAPA)

Avila FW, Goff ML, 1998. Arthropod succession patterns onto burnt carrion in two contrasting habitats in the Hawaiian Islands. Journal of Forensic Sciences, 43:581-586

Ayre GL, 1977. Exotic ants in Winnipeg. Manitoba Entomologist, 11:41-44

Baksha MW, 2001. Important pests of forest nurseries of Bangladesh and their management. Bulletin - Forest Entomology Series, Bangladesh Forest Research Institute, No.7:iii + 17 pp

Barré N, Mauleon H, Garris GI, Kermarrec A, 1991. Predators of the tick Amblyomma variegatum (Acari: Ixodidae) in Guadeloupe, French West Indies. Experimental & Applied Acarology, 12(3/4):163-170; 18 ref

Barrion AT, Bandong JP, Lumaban MD, Pantua PC, Apostol RA, Litsinger JA, 1979. Natural enemies of the rice leaf folder Cnaphalocrosis medinalis in the Philippines. International Rice Research Newsletter, 4(2):18

Bass JA, Hays SB, 1976. Geographic location and identification of fire ant species in South Carolina. Journal of the Georgia Entomological Society, 11(1):34-36

Battenfield SL, Blum MS, Battenfield SL eds, 1982. Chemistry and properties of fire ant venoms. Proceedings of the symposium on the imported fire ant, June 7-10, 1982, Atlanta American Hotel, Atlanta, Georgia, 116-118

Blum MS, 1982. Pheromonal bases of insect sociality; communications, conundrums and caveats. Les mediateurs chimiques agissant sur le comportement des insectes. Symposium international. Versailles, 16-20 novembre 1981 Institut National de la Recherche Agronomique Paris France, 149-162

Brinkman MA, Gardner WA, Ipser RM, Diffie SK, 2001. Ground-dwelling ant species attracted to four food baits in Georgia. Journal of Entomological Science, 36(4):461-463

Buren WF, 1982. Red imported fire ant now in Puerto Rico. Florida Entomologist, 65(1):188-189

Buren WF, Allen GE, Whitcomb WH, Lennartz FE, Williams RN, 1974. Zoogeography of the imported fire ants. Journal of the New York Entomological Society, 82(2):113-124

Canas LA, O'Neil RJ, 1998. Applications of sugar solutions to maize, and the impact of natural enemies on fall armyworm. International Journal of Pest Management, 44(2):59-64; 36 ref

Carroll CR, Risch SJ, 1983. Tropical annual cropping systems: ant ecology. Environmental Management, 7(1):51-57

Carroll CR, Risch SJ, 1984. The dynamics of seed harvesting in early successional communities by a tropical ant, Solenopsis geminata. Oecologia, 61(3):388-392

Castillo-Guevara C, Rico-Gray V, 2002. Is cycasin in Eumaeus minyas (Lepidoptera: Lycaenidae) a predator deterrent?. Interciencia, 27(9):465-470; many ref

Chintha RM, Das RK, Mandal SK, 2002. Pests and beneficial arthropods occurring on pointed gourd, Trichosanthes dioica Roxb. (Cucurbitaceae). Journal of Interacademicia, 6(2):174-179

Cokendolpher JC, Francke OF, 1985. Temperature preferences of four species of fire ants (Hymenoptera: Formicidae: Solenopsis). Psyche, 92(1):91-101

Collingwood CA, Tigar BJ, Agosti D, 1997. Introduced ants in the United Arab Emirates. Journal of Arid Environments, 37(3):505-512; 18 ref

Creighton WS, 1950. The ants of North America. Bulletin of the Museum of Comparative Zoolology. Harvard College, 104

Delabie JHC, Fowler HG, 1995. Soil and litter cryptic ant assemblages of Bahian cocoa plantations. Pedobiologia, 39(5):423-433

Dhiman SC, Pravesh Kumar, 2008. Biocontrol agent of Chrysocoris stolli wolf and their biocontrol efficacy. Journal of Experimental Zoology, India, 11(2):419-422

EPPO, 2014. PQR database. Paris, France: European and Mediterranean Plant Protection Organization. http://www.eppo.int/DATABASES/pqr/pqr.htm

Escalante G JA, 1979. Notes on the ants of Cusco. Revista Peruana de Entomologia, 22(1):111-112

Eskafi FM, Kolbe MM, 1990. Predation on larval and pupal Ceratitis capitata (Diptera: Tephritidae) by the ant Solenopsis geminata (Hymenoptera: Formicidae) and other predators in Guatemala. Environmental Entomology, 19(1):148-153

Essig EO, 1926. Insects of Western North America. New York, USA: Macmillan

Forel A, 1993. Formicides de l'Antille St. Vincent. Transactions of the Entomological Society of London. 41:333-4128

Fowler HG, Campiolo S, Pesquero MA, Porter SD, 1995. Notes on a southern record for Solenopsis geminata (Hymenoptera: Formicidae). Iheringia, Serie Zoologia, No. 79:173; 4 ref

Geetha Viswanathan, Ajay Narendra, 2000. The effect of urbanization on the biodiversity of ant fauna in and around Bangalore. Journal of Ecobiology, 12(2):115-122; 1 ref

Geetha Viswanathan, Paul C, Nitin Warier, Shruthi Sampath, Srimrinalini, Sushmita Jha, 2002. Neem leaf extract as a natural plant pesticide in the control of fire ants. Journal of Ecobiology, 14(2):131-135; 8 ref

George RP, Narendran TC, 1987. Ecology of Solenopsis geminate Fabr., a dominant species of ants in Malabar (India). Geobios, 14:200-204

Glancey BM, Wojcik DP, Craig CH, Mitchell JA, 1976. Ants of Mobile County, AL, as monitored by bait transects. Journal of the Georgia Entomological Society, 11(3):191-197

Heinz DJ, Carlson MK(Editors), Tabusa RS(Graphic artist), 1984. Insects. Annual report, Experiment Station, Hawaiian Sugar Planters' Association, 34-37

Hill DS, 1975. Agricultural insect pests of the tropics and their control. Agricultural insect pests of the tropics and their control. Cambridge University Press. Cambridge UK, [12 +] 516 pp

Hoffmann BD, Andersen AN, Hill GJE, 1999. Impact of an introduced ant on native rain forest invertebrates: Pheidole megacephala in monsoonal Australia. Oecologia (Berlin), 120:595-604

Hong Kong Government Information Centre, 2005. Pest-in-brief - Issue No. 5: Red Imported Fire Ants. Food and Environmental Hygiene Department, Hong Kong. http://www.fehd.gov.hk/safefood/pest-fire-ants.html

Horvitz CC, 1981. Analysis of how ant behaviours affect germination in a tropical myrmecochore Calathea microcephala (P. & E.) Koernickle (Marantaceae): Microsite selection and aril removal by neotropical ants, Odontomachus, Pachcondyla, and Solenopsis (Formicidae). Oecologia, 51:47-52

Hung ACF, Barlin MR, Vinson SB, 1977. Identification, distribution, and biology of fire ants in Texas. Publication B-1185. Texas Agricultural Experiment Station, College Station, TX

Hölldobler B, Wilson EO, 1990. The Ants. Berlin, German Federal Republic; Springer-Verlag, 732 pp

Islam Z, Way MJ, Heong KL, 2000. Methods for sampling and monitoring ants in the rice environment. Bangladesh Journal of Entomology, 10(1/2):97-114

ISSG (IUCN SSC Invasive Species Specialist Group), 2013. Global Invasive Species Database (GISD). IUCN SSC Invasive Species Specialist Group. http://www.issg.org/database/welcome/

ISSG, 2005. Global Invasive Species Database. Invasive Species Specialist Group. http://www.issg.org/database

ISSG, 2009. Global Invasive Species Database (GISD). Invasive Species Specialist Group of the IUCN Species Survival Commission. http://www.issg.org/database

Jaffe K, Villegas G, Colmenares O, Puche H, Zabala NA, Alvarez MI, Navarro JG, Pino E, 1985. Two different decision-making systems in recruitment to food in ant societies. Behaviour, 92:6-21

Kalshoven LGE, Laan PA van der (Reviser and translator), 1981. Pests of crops in Indonesia (revised). Jakarta, Indonesia: Ichtiar Baru, 701 pp

Lake PS, O'Dowd DJ, 1991. Red crabs in rain forest, Christmas Island: biotic resistance to invasion by an exotic snail. Oikos, 62(1):25-29

Lakshmikantha BP, Kumar ARV, Veeresh GK, 2001. Emergence of reproductives of the fire ant, Solenopsis geminata (Fabricius) (Hymenoptera: Formicidae), and its sex ratio. Pest Management and Economic Zoology, 9(2):129-133; 6 ref

Lakshmikantha BP, Lakshminarayan NG, Ali TMM, Veeresh GK, 1996. Fire-ant damage to potato in Bangalore. Journal of the Indian Potato Association, 23(1/2):75-76; 4 ref

Laurence GA, 1987. Pests and pest control in lawns and improved pastures. Journal of the Agricultural Society of Trinidad and Tobago, No. 87:25-29

Lee CY, Lim CY, Darah I, 2002. Survey on structure-infesting ants (Hymenoptera: Formicidae) in food preparative outlets. Tropical Biomedicine, 19:21-26

Litsinger JA, Alviola AL III, Canapi BL, 1986. Effects of flooding on insect pests and spiders in a rainfed rice environment. International Rice Research Newsletter, 11(5):24-25

Lokeshwari D, Kumar NKK, Manjunatha H, 2015. Record of ants (Hymenoptera: Formicidae) tending aphids with special reference to the melon aphid, Aphis gossypii Glover (Hemiptera: Aphididae). Pest Management in Horticultural Ecosystems, 21(1):31-37. http://aapmhe.in/index.php/pmhe/article/view/289/270

MacKay WP, Porter S, Gonzalez D, Rodriguez A, Armendedo H, Rebeles A, Vinson SB, 1990. A comparison of monogyne and polygyne populations of the tropical fire ant, Solenopsis geminata (Hymenoptera: Formicidae), in Mexico. Journal of the Kansas Entomological Society, 63(4):611-615

McInnes DA, Tschinkel WR, 1995. Queen dimorphism and reproductive strategies in the fire ant, Solenopsis geminata (Hymenoptera: Formicidae). Behavioral Ecology and Sociobiolology, 36:367-75

Moody JV, Francke OF, 1982. The ants (Hymenoptera, Formicidae) of western Texas. Part 1: Subfamily Myrmicinae. Graduate Studies, Texas Tech University, No. 27:80 pp

Moody JV, Francke OF, Merickel FW, 1981. The distribution of fire ants, Solenopsis (Solenopsis) in western Texas (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society, 54(3):469-480

Morrison LW, 1997. Polynesian ant (Hymenoptera: Formicidae) species richness and distribution: a regional survey. Acta Oecologica, 18:685-695

Morrison LW, 2000. Mechanisms of interspecific competition among an invasive and two native fire ants. Oikos, 90:238-252

Morrison LW, Kawazoe EA, Guerra R, Gilbert LE, 1999. Phenology and dispersal in Pseudacteon flies (Diptera: Phoridae), parasitoids of Solenopsis fire ants (Hymenoptera: Formicidae). Annals of the Entomological Society of America, 92(2):198-207; 41 ref

Muniappan R, Marutani M, 1992. Pest management for head cabbage production in Guam. In: Talekar S, ed. Diamondback moth and other crucifer pests. Taiwan: Asian Vegetable Research and Development Center, 541-549

Nesom GL, 1981. Ant dispersal in Wedelia hispida Hbk. (Heliantheae: Compositae). Southwestern Naturalist, 26:5-12

Nestel D, Dickschen F, 1990. The foraging kinetics of ground ant communities in different Mexican coffee agroecosystems. Oecologia, 84(1):58-63

Ota AK, Chang V, 1979. Status of control of ant damage to drip irrigation tubing. Hawaiian Sugar Technologists, 1978 Report, 79-82

Ota AK, Chang VCS, 1981. Progress in reducing ant damage to drip irrigation tubes. Hawaiian Sugar Technologists, 1980 Report, 111-112

Pavis C, Malosse C, Ducrot PH, Howse F, Jaffe K, Descoins C, 1992. Defensive secretion of first-instar larvae of rootstalk borer weevil, Diaprepes abbreviatus L. (Coleoptera: Curculionidae), to the fire-ant Solenopsis geminata (F.) (Hymenoptera: Formicidae). Journal of Chemical Ecology, 18(11):2055-2065

Perfecto I, 1991. Dynamics of Solenopsis geminata in a tropical fallow field after ploughing. Oikos, 62(2):139-144

Perfecto I, 1994. Foraging behaviour as a determinant of asymmetric competitive interaction between two ant species in a tropical agroecosystem. Oecologia, 98(2):184-192

Perfecto I, Vandermeer J, 1996. Microclimatic changes and the indirect loss of ant diversity in a tropical agroecosystem. Oecologia, 108(3):577-582; 27 ref

Porter SD, Williams DF, Patterson RS, Fowler HG, 1997. Intercontinental differences in the abundance of Solenopsis fire ants (Hymenoptera: Formicidae): escape from natural enemies. Environmental Entomology, 26(2):373-384; 40 ref

Prins AJ, 1985. Formicoidea. In: Scholtz CH, Holm E, eds. Insects of southern Africa. Durban, South Africa: Butterworths, 443-51

Risch S, 1981. Ants as important predators of rootworm eggs in the Neotropics. Journal of Economic Entomology, 74(1):88-90

Risch SJ, Carroll CR, 1982. The ecological role of ants in two Mexican agroecosystems. Oecologia, 55(1):114-119

Risch SJ, Carroll CR, 1986. Effects of seed predation by a tropical ant on competition among weeds. Ecology, USA, 67(5):1319-1327

Ross KG, Vargo EL, Fletcher DJC, 1987. Comparative biochemical genetics of three fire ant species in North America, with special reference to the two social forms of Solenopsis invicta (Hymenoptera: Formicidae). Evolution, 41(5):979-990

Sangal SK, Neena Bakshi, 1994. Feeding and foraging habits of tropical fire ant Solenopsis geminata Fabricius. Annals of Entomology, 12(1):99-102; 10 ref

Schreiner IH, Nafus DM, 1988. No-tillage and detasselling: effect on the Asian corn borer Ostrinia furnacalis and ants. Philippine Entomologist, 7(4):435-442; 17 ref

Smith MR, 1936. The ants of Puerto Rico. Journal of Agriculture, University of Puerto Rico, 20:819-75

Snelling RR, 1963. The United States species of the fire ants of the genus Solenopsis, subgenus Solenopsis Westwood with synonyms on Solenopsis aurea Wheeler. Occ. Pap. 3. California Department of Agriculture Bulletin Entomology, 15 pp

Sridhar V, Singh HS, Pandey V, 2001. Record of ant pests on Lasoda and Phalsa crops in Coastal Orissa. Insect Environment, 7(2):70; 2 ref

Suarez-Sotolongo M, 1990. Fungal disease of citrus. Memiria, Estacion Experimental de Citricos, Jaguey Grande Matanzas, Cuba, 20:72-82

Sunil Joshi, 2008. Records of aphids and aphidocolous ants (Hymenoptera: Formicidae) from Karnataka. Entomon, 33(1):15-23

Taber SW, 2000. Fire Ants. College Station, Texas: Texas A&M University Press, 308 pp

Teli VS, Salunkhe GN, 1994. Biology of sweet potato weevil. Journal of Maharashtra Agricultural Universities, 19(3):381-384; 8 ref

Tennant LE, Porter SD, 1991. Comparison of diets of two fire ant species (Hymenoptera: Formicidae): solid and liquid components. Journal of Entomological Science, 26(4):450-465; 63 ref

Torregrossa JP, Kermarrec A, 1984. Impact of two myrmecine ants (Acromyrmex landolti and Solenopsis geminata) on the development of pastures in Guiana. Prairies guyanaises et élevage bovin. Réunion interinstituts INRA, ORSTOM, GERDAT, Cayenne-Suzini, 15-16 décembre 1981., 279-290; 29 ref

Torres JA, 1984. Diversity and distribution of ant communities in Puerto Rico. Biotropica, 16:296-303

Trabanino CR, Pitre HN, Andrews KL, Meckenstock DH, 1989. Effect of seed size, colour, number of seeds per hill and depth of planting on sorghum seed survival and stand establishment: relationship to phytophagous insects. Tropical Agriculture, 66(3):225-229

Trager JC, 1991. A revision of the fire ants, Solenopsis geminata group (Hymenoptera: Formicidae: Myrmicinae). Journal of the New York Entomological Society, 99:141-198

Travis BV, 1938. The fire ant (Solenopsis spp.) As a pest of quail. Journal of Econonomic Entomology, 31:649-52

Travis BV, 1939. Tests of soil treatments for the control of the fire ant, Solenopsis geminata (F.). Journal of Economic Entomology, 32:645-50

Travis BV, 1941. Notes on the biology of the fire ant Solenopsis geminata (F.) in Florida and Georgia. Florida Entomologist, 24:15-22

Tschinkel WR, 1988. Distribution of the fire ants Solenopsis invicta and S. geminata (Hymenoptera: Formicidae) in northern Florida in relation to habitat and disturbance. Annals of the Entomological Society of America, 81(1):76-81

US Fish and Wildlife Service, 2014. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus anthracinus. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus anthracinus : US Fish and Wildlife Service.36 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0GP_I01.pdf

US Fish and Wildlife Service, 2014. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus assimulans. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus assimulans : US Fish and Wildlife Service.33 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0GQ_I01.pdf

US Fish and Wildlife Service, 2014. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus facilis. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus facilis : US Fish and Wildlife Service.32 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0GY_I01.pdf

US Fish and Wildlife Service, 2014. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus hilaris. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus hilaris : US Fish and Wildlife Service.31 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0HT_I01.pdf

US Fish and Wildlife Service, 2014. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus kuakea. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus kuakea : US Fish and Wildlife Service.29 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0VM_I01.pdf

US Fish and Wildlife Service, 2014. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus longiceps. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus longiceps : US Fish and Wildlife Service.33 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0HC_I01.pdf

US Fish and Wildlife Service, 2014. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus mana. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus mana : US Fish and Wildlife Service.30 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0VL_I01.pdf

Van den Berg H, Bagust A, Hassan K, Muhammad A, Zega S, 1995. Predation and parasitism on eggs of two pod-sucking bugs, Nezara viridula and Piezodorus hybneri, in soybean. International Journal of Pest Management, 41:134-142

Van Pelt AF, 1958. The ecology of the ants of the Welaka Reserve, Florida (Hymenoptera: Formicidae). American Midland Naturalist, 59:1-57

Vargo EL, 1993. Colony reproductive structure in a polygyne population of Solenopsis geminata (Hymenoptera: Formicidae). Annals of the Entomological Society of America, 86(4):441-449

Veeresh GK, 1990. Pest ants of India and their management. In: Veeresh GK, Mallik B, Viraklimath CA, eds. Social insects and the environment. New Delhi, India: Oxford and IBH, 267-68

Veeresh GK, 1990. Pest ants of India. In: Vander Meer RK, Jaffe K, Cedena A, eds. Applied myrmecology: A world perspective. Westview studies in insect biology. Boulder, Colorado, USA: Westview Press, 15-24

Verhaagh M, 1991. Clearing a tropical rain forest effects on the ant fauna. Tropical ecosystems: systems characteristics, utilization patterns, and conservation issues. Proceedings of the international and interdisciplinary symposium, Saarbrucken, Germany, 16-18 June 1989 [edited by Ederlen, W.; Ishwaran, N.; Muller, P.] Weirkersheim, Germany; Verlag Josef Margraf, 59-68

Vinson SB, O'Keefe S, Cook J, 2003. The common ant genera of Texas, ID guide B-6138. College Station, Texas, USA: Texas Cooperative Extension, Texas A&M University

Wauters N, Dekoninck W, Herrera HW, Fournier D, 2014. Distribution, behavioral dominance and potential impacts on endemic fauna of tropical fire ant Solenopsis geminata (Fabricius, 1804) (Hymenoptera: Formicidae: Myrmicinae) in the Galápagos archipelago. Pan-Pacific Entomologist, 90(4):205-220. http://www.bioone.org/loi/panp

Way MJ, Islam Z, Heong KL, Joshi RC, 1998. Ants in tropical irrigated rice: distribution and abundance, especially of Solenopsis geminata (Hymenoptera: Formicidae). Bulletin of Entomological Research, 88(4):467-476

Wetterer JK, O'Hara BC, 2002. Ants (Hymenoptera: Formicidae) of the Dry Tortugas, the outermost Florida Keys. Florida Entomologist, 85:303-307

Wetterer JK, Vargo DL, 2003. Ants (Hymenoptera: Formicidae) of Samoa. Pacific Science, 57(4): 409-419

Wheeler GC, Wheeler JN, 1955. The ant larvae of the myrmicine tribe Solenopsidini. American Midland Naturalist, 54:119-41

Wheeler WM, 1910. Ants: Their structure, development and behavior. New York, USA: Columbia University Press

Whitcomb WH, Denmark HA, Bhatkar AP, Greene GL, 1972. Preliminary studies on the ants of Florida soybean fields. Florida Entomologist, 55(3):129-142

Williams DF, Whelan P, 1991. Polygynous colonies of Solenopsis geminata (Hymenoptera: Formicidae) in the Galapagos Islands. Florida Entomologist, 74(2):368-371

Wilson EO, 1978. Division of labor in fire ants based on physical castes (Hymenoptera: Formicidae: Solenopsis). Journal of the Kansas Entomological Society, 51(4):615-636

Wilson EO, Brown WL, 1958. Recent changes in the introduced population of the fire ant Solenopsis saevissima (Fr. Smith). Evolution, 12:211-218

Wilson EO, Taylor RW, 1967. The ants of Polynesia (Hymenoptera: Formicidae). Pacific Insects Monograph, 14:1-109

Wojcik DP, Buren WF, Grissell EE, Carlysle T, 1976. The fire ants (Solenopsis) of Florida (Hymenoptera: Formicidae). Entomology Circular, Division of Plant Industry, Florida Department of Agriculture and Consumer Services, No. 173:4 pp

Wolcott GN, 1933. Recent experiments in the control of two Puerto Rican ants. Journal of Agriculture, University of Puerto Rico, 17:223-39

Yusa Y, 2001. Predation on eggs of the apple snail Pomacea canaliculata (Gastropoda: Ampullariidae) by the fire ant Solenopsis geminata. Journal of Molluscan Studies, 67(3):275-279

Zeng Ling, Lu YongYue, He XiaoFang, Zhang WeiQiu, Liang GuangWen, 2005. Identification of red imported fire ant, Solenopsis invicta, to invade mainland China and infestation in Wuchuan, Guangdong. Chinese Bulletin of Entomology, 42(2):144-148

Distribution References

Aesch L von, Cherix D, 2005. Introduced ant species and mechanisms of competition on Floreana Island (Galápagos, Ecuador) (Hymenoptera: Formicidae). Sociobiology. 45 (2), 463-482. http://www.csuchico.edu/biol/Sociobiology/volume/sociobiologyv45n22005.html#16

Ahmed S, Khan M R, Ahmed M, Ghaffar A, 1989. Natural enemies of paddy leaf-roller (Cnaphalocrocis medinalis Gn.). Journal of Agricultural Research (Lahore). 27 (1), 71-76.

APPPC, 1987. Insect pests of economic significance affecting major crops of the countries in Asia and the Pacific region. Technical Document No. 135., Bangkok, Thailand: Regional Office for Asia and the Pacific region (RAPA).

Avila F W, Goff M L, 1998. Arthropod succession patterns onto burnt carrion in two contrasting habitats in the Hawaiian Islands. Journal of Forensic Sciences. 43 (3), 581-586.

Ayre G L, 1977. Exotic ants in Winnipeg. Manitoba Entomologist. 41-44.

Baksha M W, 2001. Bulletin - Forest Entomology Series, Bangladesh Forest Research Institute, Chittagong, Bangladesh: Bangladesh Forest Research Institute. iii + 17 pp.

Brinkman M A, Gardner W A, Ipser R M, Diffie S K, 2001. Ground-dwelling ant species attracted to four food baits in Georgia. Journal of Entomological Science. 36 (4), 461-463.

Buren W F, 1982. Red imported fire ant now in Puerto Rico. Florida Entomologist. 65 (1), 188-189. DOI:10.2307/3494163

CABI, Undated. Compendium record. Wallingford, UK: CABI

CABI, Undated a. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI

Cañas L A, O'Neil R J, 1998. Applications of sugar solutions to maize, and the impact of natural enemies on fall armyworm. International Journal of Pest Management. 44 (2), 59-64. DOI:10.1080/096708798228329

Castillo-Guevara C, Rico-Gray V, 2002. Is cycasin in Eumaeus minyas (Lepidoptera: Lycaenidae) a predator deterrent? Interciencia. 27 (9), 465-470.

Chintha R M, Das R K, Mandal S K, 2002. Pests and beneficial arthropods occurring on pointed gourd, Trichosanthes dioica Roxb. (Cucurbitaceae). Journal of Interacademicia. 6 (2), 174-179.

Collingwood C A, Tigar B J, Agosti D, 1997. Introduced ants in the United Arab Emirates. Journal of Arid Environments. 37 (3), 505-512. DOI:10.1006/jare.1997.0309

Delabie J H C, Fowler H G, 1995. Soil and litter cryptic ant assemblages of Bahian cocoa plantations. Pedobiologia. 39 (5), 423-433.

Dhiman S C, Pravesh Kumar, 2008. Biocontrol agent of Chrysocoris stolli wolf and their biocontrol efficacy. Journal of Experimental Zoology, India. 11 (2), 419-422.

EPPO, 2014. EPPO Global database (available online). Paris, France: EPPO. https://gd.eppo.int/

EPPO, 2020. EPPO Global database. In: EPPO Global database, Paris, France: EPPO.

Escalante G J A, 1979. Notes on the ants of Cusco. (Notas sobre las hormigas del Cusco.). Revista Peruana de Entomologia. 22 (1), 111-112.

Fowler H G, Campiolo S, Pesquero M A, Porter S D, 1995. Notes on a southern record for Solenopsis geminata (Hymenoptera: Formicidae). Iheringia, Série Zoologia. 173.

George RP, Narendran TC, 1987. Ecology of Solenopsis geminate Fabr., a dominant species of ants in Malabar (India). In: Geobios, 14 200-204.

Glancey B M, Wojcik D P, Craig C H, Mitchell J A, 1976. Ants of Mobile County, AL, as monitored by bait transects. Journal of the Georgia Entomological Society. 11 (3), 191-197.

Hoffmann B D, Andersen A N, Hill G J E, 1999. Impact of an introduced ant on native rain forest invertebrates: Pheidole megacephala in monsoonal Australia. Oecologia. 120 (4), 595-604.

Hong Kong Government Information Centre, 2005. Pest-in-brief - Issue No. 5: Red Imported Fire Ants., Hong Kong: Food and Environmental Hygiene Department. http://www.fehd.gov.hk/safefood/pest-fire-ants.html

ISSG, 2005. Global Invasive Species Database., IUCN/SSC Invasive Species Specialist Group. http://www.issg.org/database

ISSG, 2009. Micropterus salmoides. In: Global Invasive Species Database (GISD), Auckland, New Zealand: University of Auckland. http://www.issg.org/database

Lake P S, O'Dowd D J, 1991. Red crabs in rain forest, Christmas Island: biotic resistance to invasion by an exotic snail. Oikos. 62 (1), 25-29. DOI:10.2307/3545442

Lakshmikantha B P, Kumar A R V, Veeresh G K, 2001. Emergence of reproductives of the fire ant, Solenopsis geminata (Fabricius) (Hymenoptera: Formicidae), and its sex ratio. Pest Management and Economic Zoology. 9 (2), 129-133.

Lakshmikantha B P, Lakshminarayan N G, Ali T M M, Veeresh G K, 1996. Fire-ant damage to potato in Bangalore. Journal of the Indian Potato Association. 23 (1/2), 75-76.

Laurence G A, 1987. Pests and pest control in lawns and improved pastures. Journal of the Agricultural Society of Trinidad and Tobago. 25-29.

Lee C Y, Lim C Y, Darah I, 2002. Survey on structure-infesting ants (Hymenoptera: Formicidae) in food preparative outlets. Tropical Biomedicine. 19 (1/2), 21-26.

Lokeshwari D, Kumar N K K, Manjunatha H, 2015. Record of ants (Hymenoptera: Formicidae) tending aphids with special reference to the melon aphid, Aphis gossypii Glover (Hemiptera: Aphididae). Pest Management in Horticultural Ecosystems. 21 (1), 31-37. http://aapmhe.in/index.php/pmhe/article/view/289/270

Morrison L W, 1997. Polynesian ant (Hymenoptera: Formicidae) species richness and distribution: a regional survey. Acta Oecologica. 18 (6), 685-695. DOI:10.1016/S1146-609X(97)80051-3

Morrison L W, Kawazoe E A, Guerra R, Gilbert L E, 1999. Phenology and dispersal in Pseudacteon flies (Diptera: Phoridae), parasitoids of Solenopsis fire ants (Hymenoptera: Formicidae). Annals of the Entomological Society of America. 92 (2), 198-207. DOI:10.1093/aesa/92.2.198

Pavis C, Malosse C, Ducrot P H, Howse F, Jaffe K, Descoins C, 1992. Defensive secretion of first-instar larvae of rootstalk borer weevil, Diaprepes abbreviatus L. (Coleoptera: Curculionidae), to the fire-ant Solenopsis geminata (F.) (Hymenoptera: Formicidae). Journal of Chemical Ecology. 18 (11), 2055-2065. DOI:10.1007/BF00981927

Perfecto I, 1991. Dynamics of Solenopsis geminata in a tropical fallow field after ploughing. Oikos. 62 (2), 139-144. DOI:10.2307/3545258

Perfecto I, Vandermeer J, 1996. Microclimatic changes and the indirect loss of ant diversity in a tropical agroecosystem. Oecologia. 108 (3), 577-582. DOI:10.1007/BF00333736

Risch S J, Carroll C R, 1986. Effects of seed predation by a tropical ant on competition among weeds. Ecology, USA. 67 (5), 1319-1327.

Ross K G, Vargo E L, Fletcher D J C, 1987. Comparative biochemical genetics of three fire ant species in North America, with special reference to the two social forms of Solenopsis invicta (Hymenoptera: Formicidae). Evolution. 41 (5), 979-990. DOI:10.2307/2409186

Schreiner I H, Nafus D M, 1988. No-tillage and detasselling: effect on the Asian corn borer Ostrinia furnacalis and ants. Philippine Entomologist. 7 (4), 435-442.

Sridhar V, Singh H S, Pandey V, 2001. Record of ant pests on Lasoda and Phalsa crops in Coastal Orissa. Insect Environment. 7 (2), 70.

Sunil Joshi, 2008. Records of aphids and aphidocolous ants (Hymenoptera: Formicidae) from Karnataka. Entomon. 33 (1), 15-23.

Teli V S, Salunkhe G N, 1994. Biology of sweet potato weevil. Journal of Maharashtra Agricultural Universities. 19 (3), 381-384.

Torregrossa J P, Kermarrec A, 1984. Impact of two myrmecine ants (Acromyrmex landolti and Solenopsis geminata) on the development of pastures in Guiana. (Incidence de deux Myrmicines (Acromyrmex landolti et Solenopsis geminata) sur le développement des pâturages en Guyane.). In: Prairies guyanaises et élevage bovin. Réunion interinstituts INRA, ORSTOM, GERDAT, Cayenne-Suzini, 15-16 décembre 1981. [Prairies guyanaises et élevage bovin. Réunion interinstituts INRA, ORSTOM, GERDAT, Cayenne-Suzini, 15-16 décembre 1981.], Paris, France: Institut National de la Récherche Agronomique. 279-290.

Torres J A, 1984. Diversity and distribution of ant communities in Puerto Rico. Biotropica. 16 (4), 296-303. DOI:10.2307/2387938

Yusa Y, 2001. Predation on eggs of the apple snail Pomacea canaliculata (Gastropoda: Ampullariidae) by the fire ant Solenopsis geminata. Journal of Molluscan Studies. 67 (3), 275-279. DOI:10.1093/mollus/67.3.275

Zeng Ling, Lu YongYue, He XiaoFang, Zhang WeiQiu, Liang GuangWen, 2005. Identification of red imported fire ant, Solenopsis invicta, to invade mainland China and infestation in Wuchuan, Guangdong. Chinese Bulletin of Entomology. 42 (2), 144-148.

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