Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide

Datasheet

Psidium guajava
(guava)

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Datasheet

Psidium guajava (guava)

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Summary

  • Last modified
  • 20 November 2019
  • Datasheet Type(s)
  • Invasive Species
  • Pest
  • Host Plant
  • Preferred Scientific Name
  • Psidium guajava
  • Preferred Common Name
  • guava
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Plantae
  •     Phylum: Spermatophyta
  •       Subphylum: Angiospermae
  •         Class: Dicotyledonae
  • Summary of Invasiveness

  • P. guajava is a fast growing tropical and subtropical species adapted to a wide range of environmental conditions. It is tolerant of shade, a precocious and prolific reproducer with seed dispersal aided by avian and mammalian vectors. It...

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Pictures

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PictureTitleCaptionCopyright
Psidium guajava; foliage and flowers.
TitleFoliage
CaptionPsidium guajava; foliage and flowers.
Copyright©A.R. Pittaway
Psidium guajava; foliage and flowers.
FoliagePsidium guajava; foliage and flowers.©A.R. Pittaway
Psidium guajava; close-up of flowers.
TitleFlowers
CaptionPsidium guajava; close-up of flowers.
Copyright©A.R. Pittaway
Psidium guajava; close-up of flowers.
FlowersPsidium guajava; close-up of flowers.©A.R. Pittaway
Psidium guajava (guava); habit.
TitleHabit
CaptionPsidium guajava (guava); habit.
Copyright©K.M. Siddiqui
Psidium guajava (guava); habit.
HabitPsidium guajava (guava); habit.©K.M. Siddiqui
Psidium guajava (guava); foliage.
TitleFoliage
CaptionPsidium guajava (guava); foliage.
Copyright©K.M. Siddiqui
Psidium guajava (guava); foliage.
FoliagePsidium guajava (guava); foliage.©K.M. Siddiqui

Identity

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Preferred Scientific Name

  • Psidium guajava L., 1753

Preferred Common Name

  • guava

Other Scientific Names

  • Guaiava pyriformis Gaertn.
  • Guajava pumila (Vahl) Kuntze
  • Guajava pyrifera (L.) Kuntze
  • Myrtus guajava (L.) Kuntze
  • Myrtus guajava var. pyrifera (L.) Kuntze
  • Psidium angustifolium Lam.
  • Psidium aromaticum Blanco
  • Psidium cujavillus Burm. f.
  • Psidium cujavus L.
  • Psidium fragrans Macfad.
  • Psidium guajava var. cujavillum (Burm.f.) Krug & Urb.
  • Psidium guajava var. minor Mattos
  • Psidium igatemyense Barb. Rodr.
  • Psidium intermedium Zipp. ex Blume
  • Psidium pomiferum L.
  • Psidium pomiferum var. sapidissimum (Jacq.) DC.
  • Psidium prostratum O. Berg
  • Psidium pumilum Vahl
  • Psidium pumilum var. guadalupense DC.
  • Psidium pyriferum L.
  • Psidium pyriferum var. glabrum Benth.
  • Psidium sapidissimum Jacq.
  • Psidium vulgare Rich.
  • Syzygium ellipticum K. Schum. & Lauterb.

International Common Names

  • English: apple guava; Brazilian guava; common guava; Guinea guava; lemon guava; pear guava; tropical guava; yellow guava
  • Spanish: guayaba; guayaba agria; guayaba blanca; guayaba cimarrona; guayaba del Perú; guayaba dulce; guayabero; guayabita; guayabita del Perú; guayabo; guyava
  • French: gouyave; goyave; goyavier; goyovier commun
  • Arabic: guwâfah
  • Chinese: fan shi liu
  • Portuguese: goiaba; goiabero; guiaiva

Local Common Names

  • Brazil: araca; araçá-goiaba; araçá-guaçú; araçá-guaiava; araçá-guiaba; araçá-uaçú; goabeira; goiaba; goiaba-branca; goiaba-pêra; goiaba-vermelha; goiabeira-branca; goibaçu; guaiaba; guava; guiava; puruí
  • Brunei Darussalam: biyabas; jambu batu
  • Cambodia: trapaek sruk
  • Cuba: guayaba cotorrera; guayabo; guayabo agrio; guayabo cotorrero; guayabo del Perú
  • Dominican Republic: guayaba injerta; guayabo
  • Germany: Guavenbaum
  • Guam: abas
  • Haiti: goavier
  • India: amarood; ettajama; jamba; jamphal; jamrukh; mansala; piyara; sapari; tupkel
  • Indonesia: jambu biji
  • Indonesia/Java: jambu klutuk
  • Italy: guaia giallo; pero del India
  • Kenya: pera
  • Laos: si da
  • Lesser Antilles: gwiyav; kwiyabu; white guava
  • Malaysia: jambu berase; jambu biji; jambu kampuchia
  • Mexico: enandi; guayaba perulera; pichi; posh
  • Myanmar: malakapen
  • Netherlands: guavaboom
  • Philippines: bayabas; guyabas
  • Saint Lucia: gwiyav
  • Tanzania: pera
  • Thailand: farang; ma-kuai; ma-man
  • USA/Hawaii: kuawa
  • Vietnam: oi

EPPO code

  • PSIGU (Psidium guajava)

Summary of Invasiveness

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P. guajava is a fast growing tropical and subtropical species adapted to a wide range of environmental conditions. It is tolerant of shade, a precocious and prolific reproducer with seed dispersal aided by avian and mammalian vectors. It can form dense thickets which displace native vegetation and is reported as an invasive weed in many countries. The balance between its valuable fruit production and its invasive potential requires careful monitoring.

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Plantae
  •         Phylum: Spermatophyta
  •             Subphylum: Angiospermae
  •                 Class: Dicotyledonae
  •                     Order: Myrtales
  •                         Family: Myrtaceae
  •                             Genus: Psidium
  •                                 Species: Psidium guajava

Notes on Taxonomy and Nomenclature

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The genus Psidium is composed of approximately 150 species of evergreen trees and shrubs in the American tropics. A good taxonomic classification of this genus is lacking. Psidium guajava is by far the most widely known and distributed (Paull and Duarte, 2012).

Psidium, except for P. guajava, is represented by di-, tetra-, hexa- and octoploid species (2n = 22) (Hirano and Nakasone, 1969). Psidium cujavillus, P. guineense and P. friedrichsthalianum from El Salvador are tetraploids (2n = 24), while the latter species from Costa Rica is a hexaploid (2n = 66). Vegetatively, plants of the two separate accessions show no visible differences (Janick and Paull, 2008).

Description

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Shallow-rooted shrub or small tree, up to 10 m tall, branching from the base and often producing suckers. Bark smooth, green to red-brown, peeling off in thin flakes. Young twigs four-angled and ridged, pubescent. Leaves opposite, with translucid punstations; petiole 3-10 mm long; blade elliptic to oblong, 5-15 x 3-7 cm, glabrous above, finely pubescent beneath, veins prominent below. Flowers solitary or in two- to three-flowered axillary cymes, about 3 cm in diameter; four to six calyx lobes, 1-1.5 cm long, irregular; petals four to five, white, 1-2 cm long; stamens numerous, 1-2 cm long; ovary 4-5-locular; style 1.5-2 cm long, stigma capitate. Fruit a berry, globose, ovoid or pyriform, 4-12 cm long, surmounted by the persistent calyx lobes; exocarp green to yellow; mesocarp fleshy, white, yellow, pink or red, sour to sweet and aromatic. Seeds numerous, yellowish, bony, reniform, 3-5 mm long, embedded in a pink or white pulp.

Plant Type

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Broadleaved
Perennial
Seed propagated
Shrub
Tree
Woody

Distribution

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The native distribution range of P. guajava is uncertain. Many botanists consider the species to be native to tropical America, probably from southern Mexico to South America, but its distribution has been greatly extended through cultivation and it is now widespread throughout the tropics and subtropics. Currently, this species is naturalized in the Old World tropics and in the West Indies (Cronk and Fuller, 1995). Some authors consider P. guajava native to Asia, perhaps due to the fact that Linnaeus in 1753 described this species based on Old World collections. However, P. guajava was reported under the name guayabo by Fernandez de Oviedo in 1535 as being widely distributed in the West Indies, both cultivated and in the wild. This is only a few decades after the discovery of the New World and therefore very unlikely that the species was introduced and was able to spread across the West Indies in such a short period of time. The species was presumably introduced into the West Indies by ancient human migration from northern South America.

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 25 Feb 2021
Continent/Country/Region Distribution Last Reported Origin First Reported Invasive Planted Reference Notes

Africa

AngolaPresentIntroduced
BotswanaPresentIntroduced
Burkina FasoPresentIntroduced
BurundiPresentIntroducedInvasive
Cabo VerdePresentIntroducedPlanted
CameroonPresentIntroducedPlanted
Central African RepublicPresentIntroduced
ComorosPresentIntroducedInvasive
Congo, Democratic Republic of thePresentIntroduced
Côte d'IvoirePresentIntroducedPlanted
EritreaPresentIntroducedPlanted
EthiopiaPresentIntroducedInvasive
GabonPresentIntroducedPlanted
GambiaPresentIntroducedPlanted
GhanaPresent
GuineaPresentIntroduced
Guinea-BissauPresentIntroduced
KenyaPresentIntroducedInvasive
MadagascarPresentIntroducedNaturalizedNaturalized
MalawiPresentIntroducedInvasive
MauritaniaPresentIntroduced
MayottePresentIntroducedInvasive
MozambiquePresentIntroduced
NamibiaPresentIntroduced
NigeriaPresentIntroducedPlanted
RéunionPresentIntroducedInvasive
RwandaPresentIntroducedInvasive
SenegalPresentIntroducedPlanted
SeychellesPresentIntroduced
South AfricaPresentIntroducedInvasivePlanted
SudanPresentIntroducedPlanted
TanzaniaPresentIntroducedInvasive
TogoPresentIntroducedPlanted
UgandaPresentIntroducedInvasive
ZambiaPresentIntroducedInvasive
ZimbabwePresentIntroducedPlantedOriginal citation: Cronk & Fuller, 1995

Asia

BangladeshPresentIntroducedPlanted
British Indian Ocean Territory
-Chagos ArchipelagoPresentIntroduced
BruneiPresentIntroducedPlanted
CambodiaPresentIntroducedPlanted
ChinaPresentIntroducedPlanted
-GuangdongPresentIntroduced
-GuangxiPresentIntroduced
-GuizhouPresentIntroduced
-HainanPresentIntroduced
-SichuanPresentIntroduced
-YunnanPresentIntroduced
Cocos IslandsPresentIntroduced
Hong KongPresentIntroducedInvasive
IndiaPresentIntroducedPlanted
-Arunachal PradeshPresent
-AssamPresentIntroduced
-ChhattisgarhPresent
-JharkhandPresent
-KeralaPresent
-OdishaPresent
-SikkimPresent
IndonesiaPresentIntroducedPlanted
-JavaPresent
-SulawesiPresentIntroduced
IsraelPresentIntroducedPlanted
JapanPresentIntroduced
LaosPresentIntroducedPlanted
MalaysiaPresentIntroducedPlanted
-Peninsular MalaysiaPresentIntroduced
MaldivesPresentIntroduced
MyanmarPresentIntroducedPlanted
PakistanPresent
PhilippinesPresentIntroducedInvasive
SingaporePresentIntroducedInvasive
Sri LankaPresentIntroducedPlanted
SyriaPresent
TaiwanPresentIntroduced
ThailandPresentIntroducedPlanted
VietnamPresentIntroducedPlanted

Europe

FrancePresent
GreecePresentIntroducedPlanted
PortugalPresentPresent based on regional distribution.
-MadeiraPresentIntroducedPlanted
SpainPresentIntroducedPlanted
-Canary IslandsPresentIntroducedPlanted

North America

AnguillaPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
Antigua and BarbudaPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
ArubaPresentIntroducedCultivated; First reported: Before 1535
BahamasPresentIntroducedInvasive
BarbadosPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
BelizePresentNative
BermudaPresentIntroducedInvasive
British Virgin IslandsPresentIntroducedNaturalizedCultivated and naturalized on Guana, Tortola, and Virgin Gorda; First reported: Before 1535
Cayman IslandsPresentIntroducedInvasive
Costa RicaPresentIntroducedInvasiveWeed in pastures
CubaPresentIntroducedInvasiveCultivated and naturalized; First reported: Before 1535
CuraçaoPresentIntroducedCultivated; First reported: Before 1535
DominicaPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
Dominican RepublicPresentIntroducedInvasiveCultivated and naturalized; First reported: Before 1535
El SalvadorPresentNative
GrenadaPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
GuadeloupePresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
GuatemalaPresentNative
HaitiPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
HondurasPresentNative
JamaicaPresentIntroducedFirst reported: Before 1535
MartiniquePresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
MexicoPresentNativePlanted
MontserratPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
Netherlands AntillesPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
NicaraguaPresentNative
PanamaPresentNative
Puerto RicoPresentIntroducedInvasiveCultivated and naturalized. Also on Mona Island, Vieques, and Culebra; First reported: Before 1535
Saint Kitts and NevisPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
Saint LuciaPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
Saint Vincent and the GrenadinesPresentIntroducedNaturalizedCultivated and naturalized; First reported: Before 1535
Trinidad and TobagoPresentIntroducedFirst reported: Before 1535
U.S. Virgin IslandsPresentIntroducedInvasiveCultivated and naturalized on St Croix, St John and St Thomas; First reported: Before 1535
United StatesPresentIntroducedPlanted
-CaliforniaPresentIntroducedPlanted
-FloridaPresentIntroduced1847Invasive
-HawaiiPresentIntroducedInvasiveFirst reported: early 1800s
-LouisianaPresentIntroduced

Oceania

American SamoaPresentIntroducedInvasive
AustraliaPresentIntroducedInvasive
-Lord Howe IslandPresentIntroducedInvasive
-New South WalesPresentIntroducedInvasive
-Northern TerritoryPresentIntroducedInvasive
-QueenslandPresentIntroducedInvasive
Christmas IslandPresentIntroducedInvasive
Cook IslandsPresentIntroducedInvasive
Federated States of MicronesiaPresentIntroducedPlanted
FijiPresentIntroducedInvasivePlanted
French PolynesiaPresentIntroducedInvasive
-Marquesas IslandsPresentIntroducedInvasivePlanted
GuamPresentIntroducedInvasive
KiribatiPresentIntroducedPlanted
Marshall IslandsPresentIntroducedPlanted
NauruPresentIntroducedPlanted
New CaledoniaPresentIntroducedInvasivePlanted
New ZealandPresentIntroducedInvasive
NiuePresentIntroducedPlanted
Norfolk IslandPresentIntroducedInvasive
Northern Mariana IslandsPresentIntroducedInvasive
PalauPresentIntroducedInvasive
Papua New GuineaPresentIntroducedPlanted
PitcairnPresentIntroducedInvasive
SamoaPresentIntroducedInvasive
Solomon IslandsPresentIntroducedInvasive
Timor-LestePresent
TongaPresentIntroducedInvasive
TuvaluPresentIntroduced
VanuatuPresentIntroducedInvasive
Wallis and FutunaPresentIntroducedInvasive

South America

ArgentinaPresentIntroducedInvasive
BoliviaPresentNative
BrazilPresentIntroducedInvasivePlanted
-AcrePresentIntroducedInvasive
-AlagoasPresentIntroduced
-AmazonasPresentIntroduced
-BahiaPresentIntroduced
-CearaPresentIntroduced
-Espirito SantoPresentIntroducedInvasive
-MaranhaoPresentIntroduced
-Mato GrossoPresentIntroduced
-Mato Grosso do SulPresentIntroducedInvasive
-Minas GeraisPresentIntroducedInvasive
-ParaibaPresentIntroducedInvasive
-ParanaPresentIntroducedInvasive
-PernambucoPresentIntroducedInvasive
-PiauiPresentIntroduced
-Rio de JaneiroPresentIntroducedInvasive
-Rio Grande do SulPresentIntroduced
-Santa CatarinaPresentIntroducedInvasive
-Sao PauloPresentIntroducedInvasive
-SergipePresentIntroduced
-TocantinsPresentIntroducedInvasive
ChilePresentPresent based on regional distribution.
-Easter IslandPresentIntroducedInvasive
ColombiaPresentNativePlanted
EcuadorPresentNative
-Galapagos IslandsPresentIntroducedInvasive
French GuianaPresentIntroducedNaturalizedCultivated and naturalized
GuyanaPresentIntroducedNaturalizedCultivated and naturalized
ParaguayPresentNative
PeruPresentNative
SurinamePresentIntroducedNaturalizedCultivated and naturalized
VenezuelaPresentNative

History of Introduction and Spread

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The Spanish took P. guajava east across the Pacific and the Portuguese took it west to Africa and India. At present it is well distributed and naturalized throughout the tropics and subtropics. Due to its ease of culture, the high nutritional value of the fruit and the popularity of the processed products, guava is important in international trade as well as in the local markets of over 60 tropical and subtropical countries. The largest producers are countries in Central and South America (Brazil, Mexico), India and Thailand (100,000 t in 1981/82), and production is increasing in the Caribbean, Hawaii, Florida (USA) and South Africa. Several authors regard P. guajava as a highly invasive species (Cronk and Fuller, 1995; Binggeli, 1999). In New Zealand it is regarded as a potential problem weed species (Owen, 1996). It was one of the first introduced species on the Galapagos islands to be recognized for its invasive behaviour (Mauchamp, 1997). It is only present on the inhabited islands and is one of a number of species that are causing serious environmental damage (Mauchamp, 1997).

In some Pacific Islands, as well as in Central American countries such as Costa Rica, P. guajava is an important weed in pastures, where it is difficult to eradicate and leads to land degradation (Somarriba, 1995). On the Pacific island of Chuuk, Space et al. (2000) list it among species that are known to be invasive elsewhere and are cultivated, common or weedy on the island. It is widely distributed on all the main Hawaiian islands, on a wide range of soil conditions, and can form dense thickets there (Smith, 1998). It is also a weed in agricultural habitats in Puerto Rico (Federal Highway Administration, 2001) and considered invasive on Bermuda. In Florida it had naturalized by 1765 (Langeland and Burks, 1998). Sheil (1994) noted that it was regenerating in disturbed areas of the Budongo forest in Uganda. Ziller and Rosa (2001) also report invasive behaviour in Brazil. Indeed, many authors link its spread to disturbance events, for example in open glades created by storms or logging activities. 

In the West Indies, P. guajava was probably introduced by aborigines before the arrival of Christopher Columbus. By 1535, Gonzalo Fernández de Oviedo, one of the first writers to chronicle the Spanish colonization of the Caribbean region, described this species as “very common everywhere in the West Indies”. This author also mentioned that native people in the West Indies cultivated and consumed the fruits of this species (Fernández de Oviedo, 1535).

 

Risk of Introduction

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This species has already been so widely introduced that it is likely to be present in almost all countries capable of growing it. The risk is therefore one of plants already present in cultivation escaping into wild environments. Given that guava production in many countries occurs in smallholdings and gardens this is likely to be difficult to control; monitoring to detect early signs of invasion would therefore be prudent. It is a category 2 invasive plant species on the Bahamas (Anon., 2003) and it appears on the Florida Exotic Pest Plant list of 1999 where it is described as a category 1 invasive that is altering plant communities (Miller et al., 2002). In South Africa it is declared a category 2 invader according to the Conservation of Agricultural Resources Act 1983 (Henderson, 2001).

Habitat

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P. guajava thrives in both humid and dry climates at altitudes of 0-1500 m (or up to 2100 m in some regions). However, the optimum yield occurs in regions with a mean temperature range of 20-30°C, and a uniform annual rainfall of 1000-2000 mm. World Agroforestry Centre (2002) describe its original habitats savannah/shrub ecotones or frequently disturbed land. In its exotic range, P. guajava is an invader of forests and forest edges, pastures and grasslands, and riparian habitats (Cronk and Fuller, 1995; Weber, 2003). In South Africa it also invades savannah and roadside habitats (Henderson, 2001) and in Florida it grows in 'hammocks, pinelands and under cypress' (Langeland and Burks, 1998).

Habitat List

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CategorySub-CategoryHabitatPresenceStatus
Terrestrial ManagedCultivated / agricultural land Present, no further details Harmful (pest or invasive)
Terrestrial ManagedManaged forests, plantations and orchards Present, no further details Harmful (pest or invasive)
Terrestrial ManagedManaged grasslands (grazing systems) Present, no further details Harmful (pest or invasive)
Terrestrial ManagedDisturbed areas Present, no further details Harmful (pest or invasive)
Terrestrial ManagedDisturbed areas Present, no further details Natural
Terrestrial ManagedRail / roadsides Present, no further details Harmful (pest or invasive)
Terrestrial Natural / Semi-naturalNatural forests Present, no further details Harmful (pest or invasive)
Terrestrial Natural / Semi-naturalNatural forests Present, no further details Natural
Terrestrial Natural / Semi-naturalRiverbanks Present, no further details Harmful (pest or invasive)
Terrestrial Natural / Semi-naturalScrub / shrublands Present, no further details Harmful (pest or invasive)
Terrestrial Natural / Semi-naturalScrub / shrublands Present, no further details Natural

Hosts/Species Affected

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P. guajava is generally an environmental weed though it is known to invade forests, both planted and natural, and managed grasslands, as well as occasionally affecting cultivated land and plantation crops.

Biology and Ecology

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Genetics

Cultivation has resulted in many varieties of this plant (Weber, 2003). In leading guava-producing countries, more or less extensive cultivar collections from all over the world are maintained, such as at the Tropical Fruit Research Station, Alstonville, Australia. Systematic breeding work was first undertaken in Hawaii to obtain better processing cultivars. Current standards for selection of processing cultivars in Hawaii include: fruit diameter at least 7.5 cm, diameter of cavity no more than 3.75 cm, fruit weight 200-300 g, seed content only 1-2%, dark pink colour, soluble solids 9-12%, vitamin C 300 mg per 100 g, flesh with few stone cells and with characteristic guava flavour. Important criteria for breeding programmes in the major fresh-fruit producing countries are yield potential, seedlessness and firm fruit that ripens slowly. Polyploidy is not uncommon in guava; parthenocarpy occurs in some diploid as well as triploid cultivars. The species shows great diversity in tree size, yield and fruit quality. Some wild seedlings produce very small seedy, gritty, musky fruit, whereas selected types can produce large, almost seedless, smooth-textured fruit with pleasant flesh (red or pink) and high flesh recovery. Dessert types have lower acidity and many are white- or yellow-fleshed. Numerous cultivars have been developed, including dual-purpose types, which are a compromise between dessert and processing requirements.

Physiology and Phenology

In good conditions, P. guajava may flower within the first 2 years. The trees reach full bearing after 5-8 years, depending on growing conditions and spacing. It is not a long-lived tree (about 40 years), but the plants may bear heavily for 15-25 years.

Reproductive Biology

Given appropriate conditions, P. guajava is able to reproduce at almost any time of year. The pollen is viable for up to 42 hours and the stigmas are receptive for about 2 days. Bees are the principal pollinators. There is some self- and cross-incompatibility. It has a hermaphrodite breeding system and produces large quantities of seeds that are then dispersed by birds and mammals. Guava seeds retain their viability for approximately 1 year at 8°C and low humidity. Vegetative reproduction from root suckers is common and impedes mechanical methods of control (Cronk and Fuller, 1995).

Environmental Requirements

P. guajava is a hardy tree that adapts to a wide range of growing conditions. In the tropics the tree is found from sea level to an altitude of about 1500 m and can stand temperatures from 15-45°C; the highest yields are recorded at mean temperatures of 23-28°C. In the subtropics, quiescent trees withstand light frosts, and 3.5-6 months of mean temperatures above 16°C (depending on the cultivar) suffice for flowering and fruiting. Guava is more drought-resistant than most tropical fruit crops. For maximum production in the tropics, however, it requires 1000-2000 mm of rain, evenly distributed over the year. If fruit ripens during a very wet period it loses flavour and may split.

Latitude/Altitude Ranges

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Latitude North (°N)Latitude South (°S)Altitude Lower (m)Altitude Upper (m)
25 -24 0 2100

Air Temperature

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Parameter Lower limit Upper limit
Absolute minimum temperature (ºC) -2 7
Mean annual temperature (ºC) 13 26
Mean maximum temperature of hottest month (ºC) 20 32
Mean minimum temperature of coldest month (ºC) 9 23

Rainfall

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ParameterLower limitUpper limitDescription
Dry season duration08number of consecutive months with <40 mm rainfall
Mean annual rainfall6003000mm; lower/upper limits

Rainfall Regime

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Bimodal
Summer
Uniform
Winter

Soil Tolerances

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Soil drainage

  • free

Soil reaction

  • acid
  • alkaline
  • neutral

Soil texture

  • heavy
  • light
  • medium

Special soil tolerances

  • infertile
  • saline

Notes on Natural Enemies

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Trees may wilt following infection by various soil fungi, and root rot caused by Phytophthora spp. is also thought to kill trees. The leaves are little affected by diseases, but anthracnose (Glomerella cingulata), which is more serious on the fruit, also affects the leaves. Blossom-end rot can become a serious disease in the rainy season; there may be physiological as well as pathological causes. Mucor rot, caused by the fungus Mucor hiemalis, spoils fruit punctured by insects. Young fruit are often seen blackened and mummified by G. cingulata or Botryodiplodia theobromae. Fruit canker, circular raised corky spots infected by Pestalotiopsis psidii, is also common. Orchard sanitation, in the form of early removal of infected plant parts, helps to reduce infections. Chemical control may be applied in the rainy season; dithane can be used against blossom-end rot and fruit canker, with additional control of anthracnose. Fruit flies are the most important pest; guava is a major host to species of Anastrepha, Ceratitis, Dacus and Argyresthia. Bagging the fruit is the main control method in continuously cropped orchards; if the crop is cycled, spraying with fenthion or use of bait sprays may provide additional means of control. Sucking insects such as scales, mealy bugs and thrips can largely be checked by predators and control of ants. Leaf-eating caterpillars and beetles may severely damage young trees; early detection greatly facilitates control by spot treatment.

Means of Movement and Dispersal

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Birds and mammals eat the fruit and disperse the seeds (Cronk and Fuller, 1995), for example rats and feral pigs (Space, 2001). According to World Agroforestry Centre (2002) the principal dispersal agents are bats. It is widely planted in gardens and as an agroforestry species and from there is able to escape cultivation and colonize disturbed sites in semi-natural and natural environments. The pantropical distribution of this species is a result of its widespread intentional introduction as a commercial fruit crop.

Impact Summary

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CategoryImpact
Animal/plant collections None
Animal/plant products None
Biodiversity (generally) Negative
Crop production Negative
Environment (generally) Negative
Fisheries / aquaculture None
Forestry production Negative
Human health None
Livestock production Negative
Native fauna None
Native flora Negative
Rare/protected species None
Tourism None
Trade/international relations None
Transport/travel None

Economic Impact

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P. guajava can cause economic impacts through the alteration of pasture habitat and because it is a host to the Caribbean fruit fly Anastrepha suspensa, a pest which can also affect citrus (Langeland and Burks, 1998). However, the economic costs of loss of pasture and costs of control must be weighed against the economic benefits accrued from the sale of fruits and processed products as commercial crops.

Environmental Impact

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This species covers large areas of the Galapagos Islands where it outcompetes native forest species (Cronk and Fuller, 1995). Similarly it is an invader of Acacia forest on Hawaii (Cronk and Fuller, 1995). Henderson (2001) classifies this species as a habitat transformer. Some authors, e.g. Smith (1998), believe it may have allelopathic effects on other plants. It is well known for forming dense thickets that can suppress the growth of native species.

Threatened Species

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Threatened SpeciesConservation StatusWhere ThreatenedMechanismReferencesNotes
Cyanea recta (Kealia cyanea)National list(s); USA ESA listing as threatened speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (1995)
Cyrtandra limahuliensis (Limahuli cyrtandra)NatureServe; USA ESA listing as threatened speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (1995)
Chrysodracon hawaiiensis (hala pepe)EN (IUCN red list: Endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (1998a)
Dubautia pauciflorula (Wahiawa Bog dubautia)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesNatureServe (2010)
Dubautia pauciflorula (Wahiawa Bog dubautia)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resources; Competition - shadingNatureServe (2010)
Isodendrion longifolium (longleaf isodendrion)EN (IUCN red list: Endangered); USA ESA listing as threatened speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2011b)
Nototrichium humile (kaala rockwort)EN (IUCN red list: Endangered); USA ESA listing as endangered speciesHawaiiCompetition - smotheringNational Tropical Botanical Garden (2007); US Fish and Wildlife Service (2008b)
Peucedanum sandwicense (makou)NatureServe; USA ESA listing as threatened speciesHawaiiCompetition - smotheringUS Fish and Wildlife Service (2011a)
Pittosporum napaliense (royal cheesewood)EN (IUCN red list: Endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resources; Competition - smotheringUS Fish and Wildlife Service (2010f)
Platydesma rostrataCR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2010g)
Poa mannii (Mann's bluegrass)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2010a)
Pritchardia napaliensisCR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2010b)
Pritchardia schattaueri (Lands of Papa pritchardia)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2009a)
Pritchardia viscosa (stickybud pritchardia)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (1998b)
Psychotria hobdyi (Hobdy's wild-coffee)USA ESA listing as endangered species; USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2010g)
Pteralyxia kauaiensis (Kauai pteralyxia)EN (IUCN red list: Endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (1995); US Fish and Wildlife Service (2010c)
Remya kauaiensis (Kauai remya)EN (IUCN red list: Endangered); USA ESA listing as endangered speciesHawaiiCompetition (unspecified)US Fish and Wildlife Service (2010d)
Scaevola coriacea (dwarf naupaka)NatureServe; USA ESA listing as endangered speciesHawaiiCompetition (unspecified)US Fish and Wildlife Service (2010g)
Schiedea apokremnos (Kauai schiedea)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition (unspecified)US Fish and Wildlife Service (2010h)
Schiedea hookeri (sprawling schiedea)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resources; Ecosystem change / habitat alterationUS Fish and Wildlife Service (2011c)
Schiedea kauaiensis (Kauai schiedea)CR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2008a)
Schiedea nuttalliiCR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (1999)
Schiedea spergulina var. leiopodaNational list(s); USA ESA listing as endangered speciesHawaiiCompetition - monopolizing resourcesUS Fish and Wildlife Service (2010i)
Sideroxylon reclinatum subsp. austrofloridense (Everglades bully)USA ESA species proposed for listingFloridaCompetition (unspecified); Ecosystem change / habitat alterationUS Fish and Wildlife Service (2013)
Urera kaalaeCR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiEcosystem change / habitat alteration; Pest and disease transmissionUS Fish and Wildlife Service (2011d)
Wilkesia hobdyi (dwarf iliau)CR (IUCN red list: Critically endangered); National list(s); USA ESA listing as endangered speciesHawaiiCompetition (unspecified)US Fish and Wildlife Service (2010e)
Xylosma crenataCR (IUCN red list: Critically endangered); USA ESA listing as endangered speciesHawaiiCompetition (unspecified)US Fish and Wildlife Service (2009b)

Social Impact

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The social impact may be considered as only positive, due to the financial and nutritional benefits from fruit production, processing and consumption.

Risk and Impact Factors

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Invasiveness
  • Proved invasive outside its native range
  • Highly adaptable to different environments
  • Tolerates, or benefits from, cultivation, browsing pressure, mutilation, fire etc
  • Highly mobile locally
  • Has high reproductive potential
  • Has propagules that can remain viable for more than one year
Impact outcomes
  • Ecosystem change/ habitat alteration
  • Negatively impacts agriculture
  • Reduced native biodiversity
Impact mechanisms
  • Competition - monopolizing resources
  • Competition - shading
  • Competition - smothering
  • Competition (unspecified)
  • Pest and disease transmission
Likelihood of entry/control
  • Highly likely to be transported internationally deliberately
  • Difficult/costly to control

Uses

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P. guajava is an ideal home garden fruit tree due to its hardiness, high yield, long supply season and high nutritive value. It is grown in orchards or incorporated into agroforestry systems in India, and is widely planted (or has spread) in Africa. Details of fruit production are given by Verheij and Coronel (1991). The potential for developing guava for a larger and wider commercial market appears to be limited, mainly by its short shelf life and its susceptibility to fruit flies. However, opportunities for expanding the processed fruit market appear to be good, in South-East Asia and in other regions. The fruit is usually eaten raw, both green and ripe (when it becomes fragrant). It is also stewed and used in shortcakes and pies. After removing the seeds, the pulp is made into preserves, jam, jelly, juice and nectar. Well-made guava jelly is deep wine-coloured, clear, of very firm consistency, and retains something of the pungent musky flavour of the fresh fruit. Guava paste (or guava cheese as it is known in the West Indies) is made by evaporating the pulp with sugar; it is eaten as a sweetmeat. The fruit, peeled, halved and cooked in light syrup, is canned and the juice and nectar are also preserved in this way. Guava powder is a good source of vitamin C and pectin. In some Asian countries the leaves are used in cooking, and medicinally against diarrhoea; they can also be used for dyeing and tanning. The wood is moderately strong and durable indoors; it is used for handles and in carpentry and turnery (Verheij and Coronel, 1991), and also for building timbers, woodware and carvings. The flowers provide nectar for bees and contribute to honey production.

Uses List

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Environmental

  • Agroforestry

Human food and beverage

  • Beverage base
  • Fruits
  • Spices and culinary herbs

Materials

  • Carved material
  • Dye/tanning
  • Wood/timber

Medicinal, pharmaceutical

  • Source of medicine/pharmaceutical
  • Traditional/folklore

Wood Products

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Sawn or hewn building timbers

  • Carpentry/joinery (exterior/interior)
  • For light construction

Woodware

  • Industrial and domestic woodware
  • Tool handles
  • Turnery

Prevention and Control

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Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.

P. guajava is able to survive some fires through regenerating vegetatively from suckers (Smith, 1998). Such vegetative reproduction from suckers is common and impedes mechanical methods of control (Cronk and Fuller, 1995). Providing all traces of the roots are removed, seedlings and young plants may be removed by pulling or digging (Weber, 2003). Cronk and Fuller (1995) report that the waxy cuticle of leaves offers some protection to the plant, reducing the effectiveness of herbicide application. If trees are felled the stumps should be treated with a herbicide to prevent regeneration through suckering (Weber, 2003). Biological control is problematic for this species because there are direct conflicts of interest with fruit growers (Smith, 1998), and none has been implemented.

Bibliography

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Amin MN, Jaiswal VS, 1988. Micropropagation as an aid to rapid cloning of a guava cultivar. Scientia Horticulturae, 36: 89-95.

Arya A, 1988. Control of Phomopsis fruit-rots of grapes and guava. Indian Phytopathology, 41(2):214-219.

Gill AS, Gangwar KS, 1990. Evaluation of intensive forage production system under guava (Psidium guajava L.) plantation. Indian Journal of Agronomy, 35(4):437-438.

Hawkes JG, 1991. The centers of plant genetic diversity in Latin America. Diversity, 7(1/2):7-9.

Kavimani S, Karpagam RI, Jaykar B, Ilango Karpagam R, 1997. Anti inflammatory activity of volatile oil of Psidium guajava. Indian Journal of Pharmaceutical Sciences, 59(3):142-144.

Menzel CM, 1985. Guava: an exotic fruit with potential in Queensland. Queensland Agricultural Journal, 111(2):93-98.

Mohyuddin AI, Piyadasa BR, Chandrasiri IMN, 1992. Integrated pest management for pests of high value crops in system B. PARC-IIBC Station, CAB International Institute of Biological Control.

Prasad NBL, Azeemoddin G, 1994. Characteristics and composition of guava (Psidium guajava L.) seed and oil. Journal of the American Oil Chemists' Society, 71(4):457.

Rehm S, Espig G, 1996. Crop plants of the tropics and subtropics. Second edition. Stuttgart, Germany: Verlag Eugen Ulmer GmbH.

Singh A, 1980. Fruit physiology and production. New Delhi, India: Kalyani Publications, 323-326.

Wilson CW III, 1980. Guava. In: Nagy S, Shaw PE, eds. Tropical and Sub tropical Fruits. Composition, Properties and Uses. Westport, Connecticut, USA: AVI Publishing Inc., 279-299.

References

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Acevedo-Rodríguez P, Strong MT, 2012. Catalogue of the Seed Plants of the West Indies. Smithsonian Contributions to Botany, 98:1192 pp. Washington DC, USA: Smithsonian Institution. http://botany.si.edu/Antilles/WestIndies/catalog.htm

Akunda EW, Oduol PA, 1989. Growth rates of some promising agroforestry tree species. Trees for development in Sub Saharan Africa. Proceedings of a regional seminar held by the International Foundation for Science (IFS), ICRAF House, Nairobi, Kenya, February 20-25, 1989, 66-70; 10 ref

Akunda EW, Oduol PA, 1989. Growth rates of some promising agroforestry tree species. Trees for development in Sub-Saharan Africa. Proceedings of a regional seminar held by the International Foundation for Science (IFS), ICRAF House, Nairobi, Kenya, 20-25 February 1989, 66-70; 10 ref

Amin MN, Jaiswal VS, 1988. Micropropagation as an aid to rapid cloning of a guava cultivar. Scientia Horticulturae, 36(1-2):89-95; 20 ref

Amin MN, Jaiswal VS, 1988. Micropropagation as an aid to rapid cloning of a guava cultivar. Scientia Horticulturae, 36(1-2):89-95; 20 ref

Anon., 2003. Invasive Plant Species. The Bahamas Environment, Science and Technology (BEST) Commission. http://www.best.bs/Invasive_plant_species.htm.

Ansar, M., Saleem, A., Iqbal, A., 1994. Cause and control of guava decline in the Punjab (Pakistan). Pakistan Journal of Phytopathology, 6(1), 41-44.

Arya A, 1988. Control of Phomopsis fruit-rots of grapes and guava. Indian Phytopathology, 41:214-219

Bajaj YPS, 1996. Biotechnology in agriculture and forestry 35. Trees IV. 1996, xix + 427 pp.; ref. at ends of chapters

Bajaj YPS, 1996. Biotechnology in agriculture and forestry 35. Trees IV. Biotechnology in agriculture and forestry 35. Trees IV., 427 pp

Beer J, 1979. Traditional agroforestry practices in the wet tropics; the 'La Suiza' case study. Activities at Turrialba, 7(3):2-5; 1 pl

Beer JW, 1979. Traditional agroforestry practices in the wet tropics; the 'La Suiza' case study. Activities at Turrialba, 7(3):2-5

Binggeli P, 1999. Invasive woody plants. http://members.lycos.co.uk/WoodyPlantEcology/invasive/index.html

Boldingh I, 1914. The flora of the Dutch West Indian Islands: The Flora of Aruba, Curaçao and Bonaire. Leyden, Netherlands: EJ. Brill, 244 pp. http://www.biodiversitylibrary.org/item/20608

Bose TK, Mitra SK, 1996. Fruits: Tropical and Subtropical. Calcutta, India: Naya Prokash

Bose TK, Mitra SK, eds, 1996. Fruits: tropical and subtropical. Calcutta, India: Naya Prokash

Broome R, Sabir K, Carrington S, 2007. Plants of the Eastern Caribbean. Online database. Barbados: University of the West Indies. http://ecflora.cavehill.uwi.edu/index.html

Buss CM, 2002. The potential threat of invasive tree species in Botswana. Department of Crop Production and Forestry, Ministry of Agriculture, Government of Botswana, 40 pp

CDFA, 2015. Guava fruit fly (GFF), https://www.cdfa.ca.gov/plant/gff/regulation.html Regulation and quarantine boundaries. California Department of Food and Agriculture. Plant Health & Pest Prevention Services

CDFA, 2015. Guava fruit fly quarantine in portion of Los Angeles County. Guava fruit fly quarantine in portion of Los Angeles County, News Release #15-025. California Department of Food and Agriculture. Office of Public Affairs

Chauhan DS, Dhyani SK, 1989. Traditional agroforestry practices in north-east Himalayan region of India. Indian Journal of Dryland Agricultural Research and Development, 4(2):73-81; 5 ref

Chauhan DS, Dhyani SK, 1989. Traditional agroforestry practices in north-east Himalayan region of India. Special issue on natural resources management for a sustainable agriculture. Indian-Journal-of-Dryland-Agricultural-Research-and-Development, 4(2):73-81; 5 ref

Cronk QCB, Fuller JL, 1995. Plant invaders: the threat to natural ecosystems. London, UK: Chapman & Hall

Cunha, R. L. da, Hubinger, M. D., Sato, A. C. K., Vieira, G. S., 2012. Tropical and subtropical fruits: postharvest physiology, processing and packaging, Iowa, USA, John Wiley & Sons.203-222.

Dadhwal KS, Sharma NK, Saroj PL, 1995. Diagnosis and possible interventions in designing agroforestry systems in north-western plains of Uttar Pradesh - a case study. Indian Journal of Soil Conservation, 23(1):47-53; 4 ref

Dadhwal KS, Sharma NK, Saroj PL, 1995. Diagnosis and possible interventions in designing agroforestry systems in north-western plains of Uttar Pradesh - a case study. Indian Journal of Soil Conservation, 23(1):47-53; 4 ref

DAISIE, 2013. Delivering Alien Invasive Species Inventories for Europe. DAISIE (online). www.europe-aliens.org

Duggan KJ, Henderson L, 1981. Progress with a survey of exotic woody plant invaders of the Transvaal. Proceedings of the fourth National Weeds Conference of South Africa., 7-20

FAO, 2013. FAOSTAT database. Rome, Italy: FAO. http://faostat3.fao.org/home/index.html [2011 data taken in 2013.]

Fernández Oviedo Gde, 1535. Historia General y Natural de las Indias: Islas y Tierra-firme del Mar Océano ([English title not available]). Madrid, Spain: Real Academia de la Historia, 770 pp

Flora of China Editorial Committee, 2012. Flora of China Web. Cambridge, USA: Harvard University Herbaria. http://flora.huh.harvard.edu/china/

Forzza RC, Leitman PM, Costa AF, Carvalho Jr AA, et al. , 2012. List of species of the Flora of Brazil (Lista de espécies Flora do Brasil). Rio de Janeiro, Brazil: Rio de Janeiro Botanic Garden. http://floradobrasil.jbrj.gov.br/2012/

Funk V, Hollowell T, Berry P, Kelloff C, Alexander SN, 2007. Checklist of the plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contributions from the United States National Herbarium, 584 pp

Gill AS, Gangwar KS, 1990. Evaluation of intensive forage production system under guava (Psidium guajava L.) plantation. Indian Journal of Agronomy, 35(4):437-438

González-Aguilar, G. A., Tiznado-Hernández, M. E., Zavaleta-Gatica, R., Martínez-Téllez, M. A., 2004. Methyl jasmonate treatments reduce chilling injury and activate the defense response of guava fruits. Biochemical and Biophysical Research Communications, 313(3), 694-701. doi: 10.1016/j.bbrc.2003.11.165

Govaerts R, 2013. World Checklist of Myrtaceae. London, UK: Royal Botanic Gardens, Kew. http://apps.kew.org/wcsp/

Hawkes JG, 1991. The centers of plant genetic diversity in Latin America. Diversity, 7(1/2):7-9

Henderson L, 2001. Alien Weeds and Invasive Plants. Plant Protection Research Institute Handbook No. 12. Cape Town, South Africa: Paarl Printers

HIRANO, R. T., NAKASONE, H. Y., 1969. Chromosome numbers of ten species and clones in the genus Psidium. Journal of the American Society of Horticultural Science, 94, 83-6.

I3N-Argentina, 2013. Base de Datos sobre Especies Invasoras (Database of invasive species)., Argentina: Universidad Nacional del Sur. www.inbiar.org.ar

I3N-Brasil, 2013. Base de dados nacional de espécies exóticas invasoras (National database of exotic invasive species). http://i3n.institutohorus.org.br/www/

Institute of Pacific Islands Forestry, 2002. Pacific Island Ecosystems at Risk (PIER) (3.3). http://www.hear.org/pier/index.html

Ito, P. J., Kunimoto, R., Ko, W. H., 1979. Transmission of mucor rot of guava fruits by three species of fruit flies. Tropical Agriculture, 56(1), 49-52.

Ito, P. J., Kunimoto, R., Ko, W., 1976. Guava cultivars tolerant to mucor fruit rot. In: 24th Annual Congress of the American Society for Horticultural Sciences, Tropical Region . 175-178.

Janick, J., Paull, R. E., 2008. The encyclopedia of fruit & nuts, CABI.xviii + 954 pp.

Jeyakumar, P., 2016. Physiological disorders in fruit crops. Guava. In: Physiological disorders in fruit crops. Guava, Tamil Nadu, India, Tamil Nadu agritech Portal.http://agritech.tnau.ac.in/agriculture/PDF/Physiological%20disorders%20-%20Dr.%20P.J.pdf

Kader, A. A., 1999. . UC Davis, Postharvest technology, Maintaining produce Quality & safety, California, USA, University of California.http://postharvest.ucdavis.edu/PFfruits/Guava/

Kavimani S, Karpagam RI, Jaykar B, Ilango Karpagam R, 1997. Anti inflammatory activity of volatile oil of Psidium guajava. Indian Journal of Pharmaceutical Sciences, 59(3):142-144

Khushk, A. M., Aslam Memon, Lashari, M. I., 2009. Factors affecting guava production in Pakistan. Journal of Agricultural Research (Lahore), 47(2), 201-210.

Kunimoto, R. K., Ito, P. J., Ko, W. H., 1977. Mucor rot of guava fruits caused by Mucor hiemalis. Tropical Agriculture, 54(2), 185-187.

Langeland KA, Burks KC, 1998. Identification and Biology of Non-native Plants in Florida’s Natural Areas. Gainesville, Florida, USA: University of Florida, 165 pp

Litz, R. E., 2004. Biotechnology of fruit and nut crops, CAB International.xxiv + 723 pp. http://www.cabi.org/cabebooks/ebook/20053001306 doi:10.1079/9780851996622.0000

Luken JO, Thieret JW, 1996. Assessment and Management of Plant Invasions. New York, USA: Springer-Verlag. 324 pp

Madagascar Catalogue, 2012. Catalogue of the Vascular Plants of Madagascar. Antananarivo, Madagascar: Missouri Botanical Garden, Madagascar Research and Conservation Program. http://www.efloras.org/madagascar

Mauchamp A, 1997. Threats from alien plant species in the Galapagos Islands. Conservation Biology, 11(1):260-263; 26 ref

Menzel CM, 1985. Guava: an exotic fruit with potential in Queensland. Queensland Agricultural Journal, 111(2):93-98

Miller JH, Chambliss EB, Bargeron CT, 2002. Invasive Plants of the Thirteen Southern States. http://www.invasive.org/seweeds.cfm

Mitchell, W. C., 1973. Hawaii Agricultural Experiment Station Miscellaneous Publication No. 111, Honolulu, Hawaii, 8-10.

Mohyuddin AI, Piyadasa BR, Chandrasiri IMN, 1992. Integrated pest management for pests of high value crops in system B. PARC-IIBC Station, CAB International Institute of Biological Control

Monzon, M. E., Biasi, B., Wang, S. J., Tang, J. M., Hallman, G. J., Mitcham, E. J., 2005. Persimmon and guava fruit response to radio frequency heating as an alternative quarantine treatment. Acta Horticulturae, (No.687), 349-350. http://www.actahort.org

Mune TL, Parham. JW, 1956. Weed control: Guava and its control in Fiji. Agricultural Journal, Fiji 27 (3/4), (103-8)

Natale, W., Mello Prado, R. de, Quaggio, J. A., Mattos, D. de Jr, 2007. In: Crisóstomo LA, Naumov A, Johnston AE, IPI Bulletin No. 18 Fertilizing for High Yield and Quality, Tropical Fruits of Brazil, Horgen, Switzerland, International Potash Institute.103-122. http://www.ipipotash.org/udocs/6_Guava.pdf

National Tropical Botanical Garden, 2007. http://ntbg.org

NatureServe, 2010. NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. Arlington, Virginia, USA: http://www.natureserve.org/explorer

Nilsson PO, 1995. Agroforestry in Cabo Delgado Mozambique: a study of farmers' experience and knowledge of growing trees. Recommendations for extension work in Cabo Delgado. Working Paper International Rural Development Centre, Swedish University of Agricultural Sciences, No. 295, v + 74 pp.; ISRN SLU-IRDC-WP-295-SE; 12 ref

Nilsson PO, 1995. Agroforestry in Cabo Delgado Mozambique: a study of farmers' experience and knowledge of growing trees. Recommendations for extension work in Cabo Delgado. Working Paper - International Rural Development Centre, Swedish University of Agricultural Sciences, No. 295:v + 74 pp.; [ISRN SLU-IRDC-WP-295-SE]; 12 ref

Owen SJ, 1996. Ecological weeds on conservation land in New Zealand: A database. Department of Conservation, Wellington, New Zealand: DOC Science Publications. http://www.hear.org/weedlists/other_areas/nz/nzecoweeds.htm

Partridge IJ, 1973. Chemical control of guava. Fiji Agricultural Journal, 35(2):101-102

Pathak RK, Ojha CM, 1993. Genetic resources of guava. Advances in horticulture: fruit crops, (1):143-147; 22 ref

Pathak RK, Ojha CM, Chadha KL (ed. ), Pareek OP, 1993. Genetic resources of guava. Advances in horticulture: fruit crops Volume 1, 143-147; 22 ref

Paull, R. E., Chen, C. C., 2014. Guava: postharvest quality-maintenance guidelines. Guava: postharvest quality-maintenance guidelines, http://www.ctahr.hawaii.edu/oc/freepubs/pdf/F_N-41.pdf Fruit, Nut and Beverage Crops F_N-41. College of Tropical Agricultural and Human Resources. Hawaii, USA: University of Hawaii.

Paull, R. E., Duarte, O., 2012. Tropical fruits, Volume 2, (Ed.2) : CABI.ix + 371 pp. http://www.cabi.org/cabebooks/ebook/20123357661 doi:10.1079/9781845937898.0000

PIER, 2012. Pacific Islands Ecosystems at Risk. Honolulu, USA: HEAR, University of Hawaii. http://www.hear.org/pier/index.html

Pleines T, 1993. Agroforestry systems and peasant farming practices for erosion control in the Antananarivo region. [Systemes agroforestiers et pratiques paysannes en lutte anti-erosive dans la region d'Antananarivo.] Akon'ny Ala, No. 11, 10-14; 11 ref

Pleines T, 1993. Agroforestry systems and peasant farming practices for erosion control in the Antananarivo region. Akon'ny Ala, 11:10-14

Prasad N, Mehta PR, Lal SB, 1952. Fusarium wilt of guava (Psidium guyava [guajava] L.) in Utter Pradesh, India. Nature, Lond. 169 (4305), (753). 7 refs

Prasad NBL, Azeemoddin G, 1994. Characteristics and composition of guava (Psidium guajava L.) seed and oil. Journal of the American Oil Chemists' Society, 71(4):457

Pritpal-Singh, Bhatt, R., Kendra-Ropar, D. P. K. V., 2013. Management of micro-nutrients deficiencies in fruit plants. In: Management of micro-nutrients deficiencies in fruit plants, Ludhiana, India, Punjab Agricultural University.http://www.agrolook.com/pdfs/micronutrients-management-plants-dec.pdf

Radha, T., Mathew, L., 2007. Fruit Crops, New Delhi, India, New India Publishing.

Raturi GB, Hiwale SS, 1993. Horti-silvi-pastoral system for increased productivity of marginal and degraded lands under rainfed conditions. Advances in Horticulture and Forestry, 3: 179-186; 3 ref

Raturi GB, Hiwale SS, 1993. Horti-silvi-pastoral system for increased productivity of marginal and degraded lands under rainfed conditions. Advances in Horticulture and Forestry, 3:179-186; 3 ref

Rehm S, Espig G, 1996. Crop plants of the tropics and subtropics. Die Kulturpflanzen der Tropen und Subtropen., Ed. 2:528 pp.; [101 fig., 55 tab.]; 2313 ref

Roxas CA, Florido HB, Saplan J, Rondina R, Dimaculangan R, 1991. Drought resistant species. RISE - Research Information Series on Ecosystems 3(2)

Saroj PL, Arora YK, 1994. Horticulture based agroforestry systems of Doon Valley. Indian Journal of Soil Conservation, 22(3):60-65; 12 ref

Saroj PL, Arora YK, 1994. Horticulture based agroforestry systems of Doon Valley. Indian Journal of Soil Conservation, 22(3):60-65; 12 ref

Sheil D, 1994. Naturalized and invasive plants in the evergreen forests of the East Usambara Mountains, Tanzania. African Journal of Ecology, 32:66-71

Singh A, 1980. Fruit physiology and production. New Delhi, India: Kalyani Publications, 323-326

Smith CW, 1998. Pest Plants of Hawaiian Native Ecosystems. University of Hawaii, USA: Department of Botany. http://www.botany.hawaii.edu/faculty/cw_smith/aliens.htm

Somarriba E (Somarriba EJ), Beer J (Beer JW), Beer JW, Somarriba EJ, 1985. Guava trees (Psidium guajava) in pastures. III. Fuelwood production. [Arboles de guayaba (Psidium guajava L.) en pastizales. III. Produccion de lena.] Turrialba, 35(4):333-338; 18 ref

Somarriba E (Somarriba EJ), Somarriba EJ, 1982. Guava (Psidium guajava L.) growing in pastures: methods of volume measurement and potential for wood production. [Guayabo (Psidium guajava L.) asociado con pastos: metodos de analisis volumetrico y potencial de produccion de lena.] 1982, iv + 33 pp.; 20 ref

Somarriba E (Somarriba EJ), Somarriba EJ, 1985. Guava trees (Psidium guajava) in pastures. I. Fruit production and seed dispersal. [Arboles de guayaba (Psidium guajava L.) en pastizales. I. Produccion de fruta y potencial de dispersion de semillas.] Turrialba, 35(3):289-295; 31 ref

Somarriba E(Somarriba EJ), 1985. Guava trees (Psidium guajava) in pastures. II. Fruit consumption and seed dispersal. Turrialba, 35(4):329-332; 9 ref

Somarriba E(Somarriba EJ), 1986. Effects of livestock on seed germination of guava (Psidium guajava L.). Agroforestry Systems, 4(3):233-238; 21 ref

Somarriba E, 1985. Guava trees in pastures. 2. Fruit consumption and seed dispersion by cows. (Arboles de guayaba (Psidium guajava L.) en pastizales. 2. Consumo de fruta y dispersion de semillas.) Turrialba, 35(4):329-332

Somarriba E, 1995. Guayaba (Psidium guajava) in pastures: establishment of live fences and rehabilitation of degraded pastures. Agroforesteri^acute~a en las Ame^acute~ricas, 2(6):27-29; 6 ref

Somarriba EJ, 1986. Effects of livestock on seed germination of guava (Psidium guajava L.). Agroforestry Systems, 4(3):233-238; 21 ref

Somarriba EJ, Beer JW, 1985. Guava trees (Psidium guajava) in pastures. III. Fuelwood production. Turrialba, 35(4):333-338; 18 ref

Space J, 2001. Notes from Jim Space on his survey of invasive plant species in Tonga. IUCN/SSC Invasive Species Specialist Group (ISSG). http://www.issg.org/features/invasives_on_tonga.html

Space JC, Waterhouse B, Denslow JS, Nelson D, Mazawa TR, 2000. Invasive plant species in Chuuk, Federated States of Micronesia. USDA Forest Service, Institute of Pacific Islands Forestry, Honolulu, Hawai'i, USA

Subramanyam MD, Iyer CPA, 1993. Improvement of guava. Advances in horticulture: fruit crops - Volume 1., 337-347; 56 ref

Subramanyam MD, Iyer CPA, Chadha KL (ed. ), Pareek OP, 1993. Improvement of guava. Advances in horticulture: fruit crops Volume 1, 337-347; 56 ref

US Fish and Wildlife Service, 1995. Recovery Plan for the Kauai Plant Cluster. In: Recovery Plan for the Kauai Plant Cluster : US Fish and Wildlife Service.270 pp.

US Fish and Wildlife Service, 1996. Big Island Plant Cluster Recovery Plan. In: Big Island Plant Cluster Recovery Plan : US Fish and Wildlife Service.202 pp.

US Fish and Wildlife Service, 1998. Big Island II: Addendum to the Recovery Plan for the Big Island Plant Cluster. In: Big Island II: Addendum to the Recovery Plan for the Big Island Plant Cluster : US Fish and Wildlife Service.80 pp. + appendices.

US Fish and Wildlife Service, 1998. Kauai II: Addendum to the Recovery Plan for the Kauai Plant Cluster. In: Kauai II: Addendum to the Recovery Plan for the Kauai Plant Cluster : US Fish and Wildlife Service.84 pp.

US Fish and Wildlife Service, 1999. Recovery Plan for Multi-Island Plants. In: Recovery Plan for Multi-Island Plants : US Fish and Wildlife Service.206 pp. + appendices.

US Fish and Wildlife Service, 2008. 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kauaiensis (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kauaiensis (no common name) : US Fish and Wildlife Service.6 pp.

US Fish and Wildlife Service, 2008. Kulu`i (Nototrichium humile). 5-Year Review. In: Kulu`i (Nototrichium humile). 5-Year Review : US Fish and Wildlife Service.11 pp.

US Fish and Wildlife Service, 2009. 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia schattaueri (Lo`ulu). In: 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia schattaueri (Lo`ulu) : US Fish and Wildlife Service.7 pp.

US Fish and Wildlife Service, 2009. 5-Year Review, Short Form Summary: Xylosma crenatum (no common name). In: 5-Year Review, Short Form Summary: Xylosma crenatum (no common name) : US Fish and Wildlife Service.7 pp.

US Fish and Wildlife Service, 2010. 5-Year Review, Short Form Summary: Species Reviewed: Poa mannii (Mann's bluegrass). In: 5-Year Review, Short Form Summary: Species Reviewed: Poa mannii (Mann's bluegrass) : US Fish and Wildlife Service.10 pp.

US Fish and Wildlife Service, 2010. 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia napaliensis (loulu palm). In: 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia napaliensis (loulu palm) : US Fish and Wildlife Service.10 pp.

US Fish and Wildlife Service, 2010. 5-Year Review, Short Form Summary: Species Reviewed: Pteralyxia kauaiensis (kaulu). In: 5-Year Review, Short Form Summary: Species Reviewed: Pteralyxia kauaiensis (kaulu) : US Fish and Wildlife Service.9 pp.

US Fish and Wildlife Service, 2010. 5-Year Review, Short Form Summary: Species Reviewed: Remya kauaiensis (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Remya kauaiensis (no common name) : US Fish and Wildlife Service.12 pp.

US Fish and Wildlife Service, 2010. 5-Year Review, Short Form Summary: Wilkesia hobdyi (dwarf iliau). In: 5-Year Review, Short Form Summary: Wilkesia hobdyi (dwarf iliau) : US Fish and Wildlife Service.8 pp.

US Fish and Wildlife Service, 2010. Determination of Endangered Status for 48 Species on Kauai and designation of Critical Habitat: Final Rule. In: Determination of Endangered Status for 48 Species on Kauai and designation of Critical Habitat: Final Rule : US Fish and Wildlife Service.i + 205 pp.

US Fish and Wildlife Service, 2010. Scaevola coriacea (dwarf naupaka). 5-Year Review: Summary and Evaluation. In: Scaevola coriacea (dwarf naupaka). 5-Year Review: Summary and Evaluation : US Fish and Wildlife Service.19 pp.

US Fish and Wildlife Service, 2010. Schiedea apokremnos (maolioli). 5-Year Review: Summary and Evaluation. In: Schiedea apokremnos (maolioli). 5-Year Review: Summary and Evaluation : US Fish and Wildlife Service.16 pp.

US Fish and Wildlife Service, 2010. Schiedea spergulina var. leiopoda (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea spergulina var. leiopoda (no common name). 5-Year Review: Summary and Evaluation : US Fish and Wildlife Service.11 pp.

US Fish and Wildlife Service, 2011. Endangered and Threatened Wildlife and Plants; Listing 23 Species on Oahu as Endangered and Designating Critical Habitat for 124 Species. In: Federal Register , 76(148) : US Fish and Wildlife Service.46362-46593. http://www.gpo.gov/fdsys/pkg/FR-2011-08-02/pdf/2011-17162.pdf

US Fish and Wildlife Service, 2011. Isodendrion longifolium (aupaka). 5-Year Review: Summary and Evaluation. In: Isodendrion longifolium (aupaka). 5-Year Review: Summary and Evaluation : US Fish and Wildlife Service.28 pp. http://ecos.fws.gov/docs/five_year_review/doc3809.pdf

US Fish and Wildlife Service, 2011. Schiedea hookeri (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea hookeri (no common name). 5-Year Review: Summary and Evaluation : US Fish and Wildlife Service.20 pp.

US Fish and Wildlife Service, 2011. Urera kaalae (opuhe). 5-Year Review: Summary and Evaluation. In: Urera kaalae (opuhe). 5-Year Review: Summary and Evaluation : US Fish and Wildlife Service.19 pp.

US Fish and Wildlife Service, 2013. U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Sideroxylon reclinatum ssp. austrofloridense. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Sideroxylon reclinatum ssp. austrofloridense : US Fish and Wildlife Service.16 pp. http://ecos.fws.gov/docs/candidate/assessments/2013/r4/Q3IM_P01.pdf

USDA-ARS, 2012. Germplasm Resources Information Network (GRIN). Online Database. Beltsville, Maryland, USA: National Germplasm Resources Laboratory. https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysearch.aspx

USDA-NRCS, 2002. The PLANTS Database, Version 3.5. National Plant Data Center, Baton Rouge, USA. http://plants.usda.gov

USDA-NRCS, 2013. The PLANTS Database. Baton Rouge, USA: National Plant Data Center. http://plants.usda.gov/

Vasquez, J., Delgado, C., Couturier, G., Ferrero, D. M., 2002. Harmful insects for the guava tree (Psidium guajava L.: Myrtaceae) in Peruvian Amazonia. Fruits (Paris), 57(5/6), 323-334. doi: 10.1051/fruits:2002029

Verheij EWM, Coronel RE(Editors), 1991. Plant resources of South-East Asia. No. 2. Edible fruits and nuts. Wageningen, Netherlands; Pudoc, 446 pp

Vieira, S. M. J., Raga, A., Benedetti, B. C., Oliveira, R. A. de, Marco, P. G. di, Scarponi, A. P. de T., 2014. Effect of ultraviolet-C radiation on "Kumagai" guavas infested by Ceratitis capitata (Diptera-Tephritidae) and on physical parameters of postharvest. Scientia Horticulturae, 165, 295-302. http://www.sciencedirect.com/science/journal/03044238 doi: 10.1016/j.scienta.2013.11.015

Villiers, E. de, Grové, T., 2000. Guava pests. Neltropika Bulletin, (No. 309), 7-10.

Vos, J. E., Schoeman, M. H., Berjak, P., Watt, M. P., Toerien, A. J., 2000. In vitro selection and commercial release of Guava Wilt resistant rootstocks. Acta Horticulturae, (No. 513), 69-79.

Weber E, 2003. Invasive plant species of the world: A reference guide to environmental weeds. Wallingford, UK: CAB International, 548 pp

Weeds of Australia, 2012. Weeds of Australia, Biosecurity Queensland Edition. http://keyserver.lucidcentral.org/weeds/data/03030800-0b07-490a-8d04-0605030c0f01/media/Html/search.html?zoom_query=

Wilson CW III, 1980. Guava. In: Nagy S, Shaw PE, eds. Tropical and Sub tropical Fruits. Composition, Properties and Uses. Westport, Connecticut, USA: AVI Publishing Inc., 279-299

Witt, A., Luke, Q., 2017. Guide to the naturalized and invasive plants of Eastern Africa, [ed. by Witt, A., Luke, Q.]. Wallingford, UK: CABI.vi + 601 pp. http://www.cabi.org/cabebooks/ebook/20173158959 doi:10.1079/9781786392145.0000

World Agroforestry Centre, 2002. Agroforestree Database. Nairobi, Kenya: ICRAF. http://www.worldagroforestrycentre.org/Sites/TreeDBS/AFT/AFT.htm

Zemede A, Ayele N, 1995. Home-gardens in Ethiopia: characteristics and plant diversity. Sinet, an Ethiopian Journal of Science, 18(2):235-266; 23 ref

Zemede A, Ayele N, 1995. Home-gardens in Ethiopia: characteristics and plant diversity. Sinet, an Ethiopian Journal of Science, 18(2):235-266; 23 ref

Ziller SR, Rosa F, 2001. Solicitaç¦o de colaboraç¦o para o workshop de 17-19 October 2001

Distribution References

Abdul-Rassoul M S, 2014. Host plants of the mealybug Nipaecoccus viridis (Newstead, 1894) (Homoptera, Pseudococcidae) in Iraq with detection of new hosts. Advances in Bio Research. 5 (4), 3-6. http://soeagra.com/abr/abrdec2014/2.pdf

Abdul-Rassoul M S, 2015. Host plants of the mealybug Nipaecoccus viridis (Newstead, 1894) (Homoptera, pseudococcidae) in Iraq with detection of new hosts. Advances in Bio Research. 6 (2), 23-26. http://soeagra.com/abr/abrmarch2015/4.pdf

Acevedo-Rodríguez P, Strong M T, 2012. Catalogue of the Seed Plants of the West Indies. Washington, DC, USA: Smithsonian Institution. 1192 pp. http://botany.si.edu/Antilles/WestIndies/catalog.htm

Adaime R, Santos R S, Azevedo T da S, Vasconcelos A da S, Sousa M do S M de, Souza Filho M F de, 2017. First record of Ceratitis capitata (Wiedemann) (Diptera: Tephritidae) in the state of Acre, Brazil. EntomoBrasilis. 10 (3), 259-260. DOI:10.12741/ebrasilis.v10i3.724

Adaime R, Sousa M do S M de, Jesus-Barros C R de, Deus E da G de, Pereira J F, Strikis P C, Souza Filho M F de, 2017a. Frugivorous flies (Diptera: Tephritidae, Lonchaeidae), their host plants, and associated parasitoids in the extreme north of Amapá State, Brazil. Florida Entomologist. 100 (2), 316-324. DOI:10.1653/024.100.0229

Adly D, Fadl H A A A, Mousa S F M, 2016. Survey and seasonal abundance of mealybug species, their parasitoids and associated predators on guava trees in Egypt. Egyptian Journal of Biological Pest Control. 26 (3), 657-664. http://www.ejbpc.com/

Akhtar N, Hanif K, Shafiq M, Anwar W, 2018. New report of Aspergillus awamori fruit rot of guava in Pakistan. JAPS, Journal of Animal and Plant Sciences. 28 (5), 1537-1541. http://www.thejaps.org.pk/docs/v-28-05/38.pdf

Ali A Y, Ahmad A M, Amar J A, Darwish R Y, Izzo A M, Al-Ahmad S A, 2016. Field parasitism levels of Ceratitis capitata larvae (Diptera: Tephritidae) by Aganaspis daci on different host fruit species in the coastal region of Tartous, Syria. Biocontrol Science and Technology. 26 (12), 1617-1625. DOI:10.1080/09583157.2016.1229756

Almeida L B M de, Coelho J B, Uchoa M A, Gisloti L J, 2019. Diversity of fruit flies (Diptera: Tephritoidea) and their host plants in a conservation unit from midwestern Brazil. Florida Entomologist. 102 (3), 562-570. DOI:10.1653/024.102.0333

Alves J C G, Brito C H de, Oliveira R de, Corsato C D A, Barbosa V de O, Silva J F da, Batista J de L, Lopes G N, 2019. Population fluctuation and faunistic analysis of fruit flies in a commercial guava orchard (Psidium guajava L.). Journal of Agricultural Science (Toronto). 11 (9), 145-153. http://www.ccsenet.org/journal/index.php/jas/article/download/0/0/39743/40675 DOI:10.5539/jas.v11n9p145

Awasthi A K, Tomar R K S, 2012. First record of the spiralling whitefly on fruit and ornamental plants in Chhattisgarh plains. Insect Environment. 18 (3/4), 72-73. http://www.currentbiotica.com/curl.aspx?url=Insect/Volume18/Insect-environment-vol-18(3)-(4).pdf

Badii K B, Billah M K, Afreh-Nuamah K, Obeng-Ofori D, 2015. Species composition and host range of fruit-infesting flies (Diptera: Tephritidae) in northern Ghana. International Journal of Tropical Insect Science. 35 (3), 137-151. http://journals.cambridge.org/action/displayJournal?jid=JTI

Basu M, Patro B, 2007. New records of host plants and natural enemies of Aphis gossypii Glover (Aphididae: Homoptera) from Orissa, India. Journal of Plant Protection and Environment. 4 (2), 74-80.

Bellis G A, Brito A A, Jesus H de, Quintao V, Sarmento J C, Bere A, Rodrigues J, Hancock D L, 2017. A preliminary account of the fruit fly fauna of Timor-Leste (Diptera: Tephritidae: Dacinae). Zootaxa. 4362 (3), 421-432. http://www.mapress.com/j/zt/article/view/zootaxa.4362.3.6 DOI:10.11646/zootaxa.4362.3.6

Boldingh I, 1914. The flora of the Dutch West Indian Islands: The Flora of Aruba, Curaçao and Bonaire., Leyden, Netherlands: EJ Brill. 244 pp. http://www.biodiversitylibrary.org/item/20608

Brhane D, Girmay M, Debru H, Hayelom M, Tekie M H M, Sharma V K, Naqvi S D Y, 2018. Survey on insect-pests of guava and management practices around Hamelmalo sub-zoba. Journal of Eco-friendly Agriculture. 13 (1), 69-71. http://ecoagrijournal.com/wp-content/uploads/2019/10/Full-paper131.pdf

Broome R, Sabir K, Carrington S, 2007. Plants of the Eastern Caribbean. Online database., Barbados: University of the West Indies. http://ecflora.cavehill.uwi.edu/index.htm

Buss CM, 2002. The potential threat of invasive tree species in Botswana., Department of Crop Production and Forestry, Ministry of Agriculture, Government of Botswana. 40 pp.

Caasi-Lit M T, Lit I L Jr, Larona A R, 2012. Expansion of local geographic and host ranges of Nipaecoccus nipae (Maskell) (Pseudococcidae, Hemiptera) in the Philippines with new records of predators and attending ants. Philippine Journal of Crop Science. 37 (1), 47-56.

CABI, Undated. Compendium record. Wallingford, UK: CABI

CABI, Undated a. CABI Compendium: Status inferred from regional distribution. Wallingford, UK: CABI

CABI, Undated b. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI

Carranza M, Larran S, Ronco B, 2002. First report of Glomerella cingulata on common guava in Argentina. Plant Disease. 86 (4), 440. DOI:10.1094/PDIS.2002.86.4.440B

DAISIE, 2013. Delivering Alien Invasive Species Inventories for Europe. http://www.europe-aliens.org/

Danggomen A, Visarathanonth N, Manoch L, Piasai O, 2013. Morphological studies of endophytic and plant pathogenic Phomopsis liquidambaris and Diaporthe phaseolorum (P. phaseoli anamorph) from healthy plants and diseased fruits. Thai Journal of Agricultural Science. 46 (3), 157-164. http://www.thaiagj.org/images/stories/Journal_online/2013/3/07-tj-agr-1013-66-p157-164.pdf

Debabrata Sen, 2015. Some species of Dorylaimoidea (Nematoda) associated with guava and litchi, with new records from West Bengal, India. Records of the Zoological Survey of India. 115 (1), 7-19.

Etienne J, Matile-Ferrero D, Kondo T, 2014. Phalacrococcus howertoni Hodges & Hodgson (Hemiptera: Coccoidea: Coccidae), a new soft scale record for the island of Guadeloupe. Revista Corpoica - Ciencia y Tecnologia Agropecuarias. 15 (1), 115-118. http://www.corpoica.org.co/SitioWeb/Archivos/Revista/Articulo10_REVISTACORPOICAVol15No1-2014.pdf

Fadlelmula A A, Ali E B M, 2014. Fruit fly species, their distribution, host range and seasonal abundance in Blue Nile State, Sudan. Persian Gulf Crop Protection. 3 (3), 17-24. http://www.cropprotection.ir/files_site/paperlist/Journal3-3-140928102059.pdf

FAO, 2013. FAOSTAT database., Rome, Italy: FAO. http://faostat3.fao.org/home/index.html

Flora of China Editorial Committee, 2012. Flora of China Web., Cambridge, USA: Harvard University Herbaria. http://flora.huh.harvard.edu/china/

Forzza RC, Leitman PM, Costa AF, Carvalho Jr AA et al, 2012. List of species of the Flora of Brazil. (Lista de espécies Flora do Brasil)., Rio de Janeiro, Brazil: Rio de Janeiro Botanic Garden. http://floradobrasil.jbrj.gov.br/2012/

Funk V, Hollowell T, Berry P, Kelloff C, Alexander S N, 2007. Contributions from the United States National Herbarium, Washington, USA: Department of Systematic Biology - Botany, National Museum of Natural History, Smithsonian Institution. 55, 584 pp.

Galanihe L D, Jayasundera M U P, Vithana A, Asselaarachchi N, Watson G W, 2010. Occurrence, distribution and control of papaya mealybug, Paracoccus marginatus (Hemiptera: Pseudococcidae), an invasive alien pest in Sri Lanka. Tropical Agricultural Research and Extension. 13 (3), 81-86. http://www.sljol.info/index.php/TARE/article/view/3143/2522

Ghirtlahre S K, Awasthi A K, Ganguli J L, 2014. New record of spiraling white fly, Aleurodicus dispersus Russell (Hemiptera: Aleyrodidae), on mistletoe, Dendrophthoe falcata (L.f) Ettingsh. Insect Environment. 20 (3), 80. http://www.currentbiotica.com/Insect/Volume20-3/IE-20(3)-Paper-5.pdf

Gonçalves G B, Santos J C G dos, Silva C E da, Santos E S dos, Nascimento R R do, Sant'ana A E G, Zucchi R A, 2006. Occurrence of fruit flies (Diptera: Tephritidae) in the state of Alagoas, Brazil. Florida Entomologist. 89 (1), 93-94. DOI:10.1653/0015-4040(2006)89[93:OOFFDT]2.0.CO;2

Govaerts R, 2013. World Checklist of Myrtaceae., London, UK: Royal Botanic Gardens, Kew. http://apps.kew.org/wcsp/

Hassani I M, Raveloson-Ravaomanarivo L H, Delatte H, Chiroleu F, Allibert A, Nouhou S, Quilici S, Duyck P F, 2016. Invasion by Bactrocera dorsalis and niche partitioning among tephritid species in Comoros. Bulletin of Entomological Research. 106 (6), 749-758. DOI:10.1017/S0007485316000456

Hussain M A, Haile A, Ahmad T, 2015. Infestation of two tephritid fruit flies, Bactrocera dorsalis (syn. B. invadens) and Ceratitis capitata, in guava fruits from selected regions of Eritrea. African Entomology. 23 (2), 510-513. DOI:10.4001/003.023.0214

I3N-Argentina, 2013. Database of invasive species. (Base de Datos sobre Especies Invasoras)., Argentina: Universidad Nacional del Sur. http://www.inbiar.org.ar

I3N-Brasil, 2013. National database of exotic invasive species. (Base de dados nacional de espécies exóticas invasoras)., http://i3n.institutohorus.org.br/www/

Ibrahim I K A, Handoo Z A, 2016. Occurrence of phytoparasitic nematodes on some crop plants in northern Egypt. Pakistan Journal of Nematology. 34 (2), 163-169.

Ibrahim I K A, Mokbel A A, Handoo Z A, 2010. Current status of phytoparasitic nematodes and their host plants in Egypt. Nematropica. 40 (2), 239-262. http://www.ontaweb.org/

Institute of Pacific Islands Forestry, 2002. Pacific Island Ecosystems at Risk (PIER)., http://www.hear.org/pier/index.html

Iwahori H, Truc N T N, Ban D V, Ichinose K, 2009. First report of root-knot nematode Meloidogyne enterolobii on guava in Vietnam. Plant Disease. 93 (6), 675. http://apsjournals.apsnet.org/loi/pdis DOI:10.1094/PDIS-93-6-0675C

Jayanthi P D K, Nagaraja T, Raghava T, Vivek Kempraj, 2016. Pomegranate, a newly documented host plant of tea mosquito bug, Helopeltis antonii Signoret. Pest Management in Horticultural Ecosystems. 22 (1), 88-90. http://aapmhe.in/index.php/pmhe/article/view/711/664

Kanmiya K, Sonobe R, 2002. Records of two citrus pest whiteflies in Japan with special reference to their mating sounds (Homoptera: Aleyrodidae). Applied Entomology and Zoology. 37 (3), 487-495. DOI:10.1303/aez.2002.487

Katiyar K P, Camacho Molina J, Matheus R, 2000. Fruit flies (Diptera: Tephritidae) infesting fruits of the genus Psidium (Myrtaceae) and their altitudinal distribution in western Venezuela. Florida Entomologist. 83 (4), 480-486. DOI:10.2307/3496724

Khan M R, Hassan A, Ghosh B, Das B, Ghosh S, Ray S K, 2007. Diversity and community analyses of soil nematodes associated with guava from West Bengal, India. Acta Horticulturae. 483-487. http://www.actahort.org

Khushk A M, Aslam Memon, Lashari M I, 2009. Factors affecting guava production in Pakistan. Journal of Agricultural Research (Lahore). 47 (2), 201-210.

Kolla Sreedevi, Reddy P V R, Sandeep Singh, Badal Bhattacharyya, Bhagawati S, 2019. White grubs (Coleoptera: Scarabaeidae) on fruit crops: emerging as pests of economic importance. Pest Management in Horticultural Ecosystems. 25 (2), 263-267. http://www.aapmhe.in/index.php/pmhe/article/view/929/811

Kondo T, Gullan P J, Peronti A L B G, Ramos-Portilla A A, Caballero A, Villarreal-Pretelt N, 2016. First records of the iceryine scale insects Crypticerya brasiliensis (Hempel) and Crypticerya genistae (Hempel) (Hemiptera: Monophlebidae) for Colombia. Insecta Mundi. 1-9. http://centerforsystematicentomology.org/insectamundi/PDF-download.asp?FileName=0480_Kondo_etal_2016.pdf

Kumar A, 2013. Fig wax scale, Ceroplastes rusci an emerging pest of Dalbergia sissoo and its parasitisation in India. International Journal of Current Science. 106-114. http://www.currentsciencejournal.info/issuespdf/Dr%20Arvind16.pdf

Kumar V, Seal D R, Kakkar G, McKenzie C L, Osborne L S, 2012. New tropical fruit hosts of Scirtothrips dorsalis (Thysanoptera: Thripidae) and its relative abundance on them in South Florida. Florida Entomologist. 95 (1), 205-207. DOI:10.1653/024.095.0134

Kumawat M M, Singh K M, Wangchu L, 2017. First report of an invasive longhorn beetle, Aristobia reticulator (Voet) (Coleoptera: Cerambycidae) in litchi, Litichi chinensis Sonn. (Sapindaceae), in India. Coleopterists Bulletin. 71 (1), 131-136. DOI:10.1649/0010-065X-71.1.131

Lasa R, Tadeo E, 2015. Invasive drosophilid pests Drosophila suzukii and Zaprionus indianus (Diptera: Drosophilidae) in Veracruz, Mexico. Florida Entomologist. 98 (3), 987-988. http://www.bioone.org/loi/flen DOI:10.1653/024.098.0332

Lemos L do N, Adaime R, Jesus-Barros C R de, Deus E da G de, 2014. New hosts of Bactrocera carambolae (Diptera: Tephritidae) in Brazil. Florida Entomologist. 97 (2), 841-843. DOI:10.1653/024.097.0274

Lemos L do N, Deus E da G de, Nascimento D B do, Jesus-Barros C R de, Costa Neto S V da, Adaime R, 2017. Species of Anastrepha (Diptera: Tephritidae), their host plants, and parasitoids in small fruit production areas in the state of Amapá, Brazil. Florida Entomologist. 100 (2), 403-410. DOI:10.1653/024.100.0201

Lokeshwari D, Kumar N K K, Manjunatha H, 2015. Molecular diversity of the Aphis gossypii (Hemiptera: Aphididae): a potential vector of potyviruses (Potyviridae). Annals of the Entomological Society of America. 108 (4), 621-633. DOI:10.1093/aesa/sav034

Madagascar Catalogue, 2012. Catalogue of the Vascular Plants of Madagascar., Antananarivo, Madagascar: Missouri Botanical Garden, Madagascar Research and Conservation Program. http://www.efloras.org/madagascar

Mala B R J, Raghava T, Jayanthi P D K, 2019. Incidence of wax scale, genus Ceroplastes sp. nr. pseudoceriferus (Hemiptera: Coccidae) on guava (Psidium guajava L.). Pest Management in Horticultural Ecosystems. 25 (2), 268-269. http://www.aapmhe.in/index.php/pmhe/article/view/922/813

Mastoi M I, Azura A N, Muhamad R, Idris A B, Ibrahim Y, 2014. Survey of papaya mealybug, Paracoccus marginatus (Hemiptera: Pseudococcidae) and its natural enemies in Penninsular Malaysia. Pakistan Journal of Agriculture, Agricultural Engineering, Veterinary Sciences. 30 (2), 172-186. http://www.sau.edu.pk/sau_journal/2015/vol-30-2015/Paper-6%20vol.30(2)%20Dec.2014.pdf

Mauchamp A, 1997. Threats from alien plant species in the Galápagos Islands. Conservation Biology. 11 (1), 260-263. DOI:10.1046/j.1523-1739.1997.95356.x

Meena Thakur, Dinabandhoo C L, 2005. Predatory mites associated with phytophagous mites of temperate and sub-tropical fruit trees in Himachal Pradesh. Journal of Biological Control. 19 (1), 81-84.

Mware B, Olubayo F, Narla R, Songa J, Amata R, Kyamanywa S, Ateka E M, 2010. First record of spiraling whitefly in coastal Kenya: emergence, host range, distribution and association with cassava brown streak virus disease. International Journal of Agriculture and Biology. 12 (3), 411-415. http://www.fspublishers.org/ijab/past-issues/IJABVOL_12_NO_3/18.pdf

Mwatawala M W, Meyer M de, Makundi R H, Maerere A P, 2009. Host range and distribution of fruit-infesting pestiferous fruit flies (Diptera, Tephritidae) in selected areas of Central Tanzania. Bulletin of Entomological Research. 99 (6), 629-641. DOI:10.1017/S0007485309006695

Ocampo Flórez V, Durán Prieto J, Albornoz M, Forero D, 2018. New plant associations for Monalonion velezangeli (Hemiptera: Miridae) in green urban areas of Bogotá (Colombia). Acta Biológica Colombiana. 23 (2), 205-208. https://revistas.unal.edu.co/index.php/actabiol/article/view/69374/pdf_art8

Owen S J, 1996. Ecological weeds on conservation land in New Zealand: a database. In: Ecological weeds on conservation land in New Zealand: a database, Department of Conservation, Wellington, New Zealand: DOC Science Publications. http://www.hear.org/weedlists/other_areas/nz/nzecoweeds.htm

PIER, 2012. Pacific Islands Ecosystems at Risk., Honolulu, USA: HEAR, University of Hawaii. http://www.hear.org/pier/index.html

Poornima K, Suresh P, Kalaiarasan P, Subramanian S, Ramaraju K, 2016. Root knot nematode, Meloidogyne enterolobii in guava (Psidium guajava L.) a new record from India. Madras Agricultural Journal. 103 (10/12), 359-365. https://drive.google.com/open?id=0B20TM0cqzsQmT0VtYWpOQ2RvMEE

Prabhakar C S, Pankaj Sood, Mehta P K, 2012. Fruit fly (Diptera: Tephritidae) diversity in cucurbit fields and surrounding forest areas of Himachal Pradesh, a north-western Himalayan state of India. Archives of Phytopathology and Plant Protection. 45 (10), 1210-1217. http://www.tandfonline.com/loi/gapp20 DOI:10.1080/03235408.2012.660612

Quénéhervé P, Godefroid M, Mège P, Marie-Luce S, 2011. Diversity of Meloidogyne spp. parasitizing plants in Martinique Island, French West Indies. Nematropica. 41 (2), 191-199. http://journals.fcla.edu/nematropica/article/view/76630/74240

Rashtra Vardhana, 2017. Plant's diseases of district Ghaziabad and adjacent areas. Plant Archives. 17 (1), 727-732. http://www.plantarchives.org/PDF%2017-1/727-732%20(3511).pdf

Renu Srivastava, Lal A A, 2009. Incidence of post-harvest fungal pathogens in guava and banana in Allahabad. Journal of Horticultural Sciences. 4 (1), 85-89.

Romo J P, Osorio J G M, Yepes M S, 2012. Identification of new hosts for Ralstonia solanacearum (Smith) race 2 from Colombia. Revista de Protección Vegetal. 27 (3), 151-161. http://scielo.sld.cu/scielo.php?script=sci_arttext&pid=S1010-27522012000300003&lng=en&nrm=iso&tlng=en

Sakthivel P, Karuppuchamy P, Kalyanasundaram M, Srinivasan T, 2012. Host plants of invasive papaya mealybug, Paracoccus marginatus (Williams and Granara de Willink) in Tamil Nadu. Madras Agricultural Journal. 99 (7/9), 615-619. https://doc-00-7g-docsviewer.googleusercontent.com/viewer/securedownload/dsn1aovipa7l846lsfcf94nedj8q2p4u/qo3phtufamvk9q39umu888pbj4t4kkc6/1348647300000/c2l0ZXM=/AGZ5hq8BgbJY1gwaOYx83cPOdNw6/WkdWbVlYVnNkR1J2YldGcGJud3hNWFJvWlcxaFpISmhjMkZuY21samRXeDBkWEpoYkdwdmRYSnVZV3g4WjNnNk56WmpPREk1WXpBd01XWTNZelZrWkE=?a=gp&filename=99-7-9-615-619.pdf&chan=EQAAAOqeu1nfMdjbyOfMSElqQCfRbAOx1kCMBqnRUfeLUnjy&docid=0508176bd4abbdc3e7017b1a89751bc3%7C9c9df36583445f1fe402a841b5e1963b&sec=AHSqidZmGWqJKVKwfKsaqtFstCH

Salazar Yepes M, Carvalho Júnior A A de, 2013. Uredinales (rust fungi) biota of the Parque Nacional do Itatiaia, Brazil: an analysis of composition, species diversity and altitudinal distribution. Caldasia. 35 (1), 165-176. http://www.scielo.org.co/scielo.php?script=sci_arttext&pid=S0366-52322013000100012&lng=en&nrm=iso&tlng=en

Sanahuja G, Lopez P, Palmateer A J, Chase A R, 2018. Red rust of Neoregelia bromeliads caused by a parasitic alga Cephaleuros parasiticus in Florida. Plant Health Progress. PHP-11-17-0068-RS. http://www.plantmanagementnetwork.org/php/elements/sum2.aspx?id=11008

Santos O O, Castellani M A, Bittencourt M A L, Moreira A A, Strikis P C, 2018. Frugivorous flies (Diptera: Lonchaeidae) hosts in the state of Bahia, Brazil and registers of new bitrophic interactions. Brazilian Journal of Biology. 78 (3), 591-592. http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1519-69842018000300591&lng=en&nrm=iso&tlng=en

Sartiami D, Mound L A, 2013. Identification of the terebrantian thrips (Insecta, Thysanoptera) associated with cultivated plants in Java, Indonesia. ZooKeys. 1-21. http://www.pensoft.net/journals/zookeys/article/5455/identification-of-the-terebrantian-thrips-insecta-thysanoptera-associated-with-cultivated-plants-in-java-indonesia

Schliserman P, Ovruski S, Colin C, Norrbom A L, Aluja M, 2004. First report of Juglans australis (Juglandaceae) as a natural host plant for Anastrepha schultzi (Diptera: Tephritidae) with notes on probable parasitism by Doryctobracon areolatus, D. brasiliensis, Opius bellus (Braconidae) and Aganaspis pelleranoi (Figitidae). Florida Entomologist. 87 (4), 597-599. DOI:10.1653/0015-4040(2004)087[0597:FROJAJ]2.0.CO;2

Selvaraj K, Ankita Gupta, Venkatesan T, Jalali S K, Sundararaj R, Ballal C R, 2017. First record of invasive rugose spiraling whitefly Aleurodicus rugioperculatus Martin (Hemiptera: Aleyrodidae) along with parasitoids in Karnataka. Journal of Biological Control. 31 (2), 74-78. http://informaticsjournals.com/index.php/jbc/article/viewFile/16015/15196 DOI:10.18311/jbc/2017/16015

Shahina F, Erum Y I, 2007. Distribution of cyst nematodes in Pakistan. Pakistan Journal of Nematology. 25 (1), 29-35.

Shamesa Maryam, Sandhu A A, Imran Bodlah, Aziz M A, Ayesha Aihetasham, 2019. Contribution to Aphid's fauna of Gujranwala (Punjab), Pakistan. Punjab University Journal of Zoology. 34 (1), 9-16. http://pu.edu.pk/images/journal/zology/PDF-FILES/2-34_1_19.pdf

Sheil D, 1994. Naturalized and invasive plant species in the evergreen forests of the East Usambara Mountains, Tanzania. African Journal of Ecology. 32 (1), 66-71. DOI:10.1111/j.1365-2028.1994.tb00556.x

Silva M do C L da, Santos C D G, 2017. Distribution of Meloidogyne enterolobii in guava orchards in the state of Ceará, Brazil. Revista Caatinga. 30 (2), 335-342. https://periodicos.ufersa.edu.br/revistas/index.php/caatinga/article/view/4754/pdf

Sirisena U G A I, Watson G W, Hemachandra K S, Wijayagunasekara H N P, 2013. Mealybugs (Hemiptera: Pseudococcidae) species on economically important fruit crops in Sri Lanka. Tropical Agricultural Research. 25 (1), 69-82. http://www.pgia.ac.lk/sites/default/files/congress/journel/v25/Journal-No%201/Papers/7%20U.G.A.I.%20Sirisena_final_final_edited.pdf

Sunil Joshi, Sangma R H Ch, 2015. Natural enemies associated with aphids and coccids from Sikkim, India. Journal of Biological Control. 29 (1), 3-7. http://journalofbiologicalcontrol.com/index.php/jbc/article/view/75778/59026

Sunpapao A, Thithuan N, Bunjongsiri P, Arikit S, 2016. Cephaleuros parasiticus, associated with algal spot disease on Psidium guajava in Thailand. Australasian Plant Disease Notes. 11 (1), 12. http://link.springer.com/article/10.1007/s13314-016-0199-0

Taira T L, Abot A R, Nicácio J, Uchôa M A, Rodrigues S R, Guimarães J A, 2013. Fruit flies (Diptera, Tephritidae) and their parasitoids on cultivated and wild hosts in the Cerrado-Pantanal ecotone in Mato Grosso do Sul, Brazil. Revista Brasileira de Entomologia. 57 (3), 300-308. HTTP://www.scielo.br/rbent DOI:10.1590/S0085-56262013000300007

Tanga C M, Ekesi S, Govender P, Mohamed S A, 2016. Host-plant relationships and natural enemies of the invasive mealybug, Rastrococcus iceryoides Green in Kenya and Tanzania. Journal of Applied Entomology. 140 (9), 655-668. http://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1439-0418 DOI:10.1111/jen.12292

Trindade T D, Trevisan H, Carvalho A G de, 2019. Incidence of Bemisia tabaci Gennadius (Aleyrodidae, aleyrodinae) in four Atlantic forest fragments. Arquivos do Instituto Biológico (São Paulo). DOI:10.1590/1808-1657000942017

USDA-ARS, 2012. Germplasm Resources Information Network (GRIN). Online Database. Beltsville, Maryland, USA: National Germplasm Resources Laboratory. https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysimple.aspx

USDA-NRCS, 2002. The PLANTS Database. Greensboro, North Carolina, USA: National Plant Data Team. https://plants.sc.egov.usda.gov

USDA-NRCS, 2013. The PLANTS Database. Greensboro, North Carolina, USA: National Plant Data Team. https://plants.sc.egov.usda.gov

Vargas-Madriz H, Martínez-Damian M T, Lázaro-Dzul M O, Terán-Vargas A P, Azuara-Domínguez A, 2016. New distribution registry of guava weevil, Conotrachelus dimidiatus Champion, in guava (Psidium guajava L.) in Mexico. Southwestern Entomologist. 41 (3), 883-885. DOI:10.3958/059.041.0332

Verma S P, Dinabandhoo C L, 2005. Armoured scales (Homoptera: Diaspididae) associated with temperate and subtropical fruit trees in Himachal Pradesh. Acta Horticulturae. 423-426. http://www.actahort.org

Vieira F N da S, Sousa E M de, Louzeiro L R F, Braga e Silva S, 2019. Lonchaeidae (Diptera) species and their host plants in the Cerrado biome in the state of Piauí, Brazil. Arquivos do Instituto Biológico (São Paulo). DOI:10.1590/1808-1657000242018

Weber E, 2003. Invasive plant species of the world: a reference guide to environmental weeds. [ed. by Weber E]. Wallingford, UK: CABI Publishing. viii + 548 pp.

Weeds of Australia, 2012. Weeds of Australia, Biosecurity Queensland Edition., http://keyserver.lucidcentral.org/weeds/data/03030800-0b07-490a-8d04-0605030c0f01/media/Html/search.html?zoom_query=

Witt A, Beale T, Wilgen B W van, 2018. An assessment of the distribution and potential ecological impacts of invasive alien plant species in eastern Africa. Transactions of the Royal Society of South Africa. 73 (3), 217-236. DOI:10.1080/0035919X.2018.1529003

Witt A, Luke Q, 2017. Guide to the naturalized and invasive plants of Eastern Africa. [ed. by Witt A, Luke Q]. Wallingford, UK: CABI. vi + 601 pp. http://www.cabi.org/cabebooks/ebook/20173158959 DOI:10.1079/9781786392145.0000

World Agroforestry Centre, 2002. Agroforestree Database., Nairobi, Kenya: ICRAF. http://www.worldagroforestrycentre.org/Sites/TreeDBS/AFT/AFT.htm

Yugendra K, Gopali J B, Athani S I, Tulsiram K, 2020. Survey and surveillance of fruit fly on guava across Karnataka. Journal of Experimental Zoology, India. 23 (1), 485-489. http://www.connectjournals.com/jez

Ziller SR, Rosa F, 2001. (Solicitação de colaboração para o workshop de 17-19 October 2001).,

Links to Websites

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WebsiteURLComment
GISD/IASPMR: Invasive Alien Species Pathway Management Resource and DAISIE European Invasive Alien Species Gatewayhttps://doi.org/10.5061/dryad.m93f6Data source for updated system data added to species habitat list.
Global register of Introduced and Invasive species (GRIIS)http://griis.org/Data source for updated system data added to species habitat list.

Contributors

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01/03/13 Updated by:

Julissa Rojas-Sandoval, Department of Botany-Smithsonian NMNH, Washington DC, USA

Pedro Acevedo-Rodríguez, Department of Botany-Smithsonian NMNH, Washington DC, USA

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