Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide

Datasheet

Saccharicoccus sacchari
(grey sugarcane mealybug)

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Datasheet

Saccharicoccus sacchari (grey sugarcane mealybug)

Summary

  • Last modified
  • 29 March 2018
  • Datasheet Type(s)
  • Pest
  • Preferred Scientific Name
  • Saccharicoccus sacchari
  • Preferred Common Name
  • grey sugarcane mealybug
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Metazoa
  •     Phylum: Arthropoda
  •       Subphylum: Uniramia
  •         Class: Insecta

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Pictures

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PictureTitleCaptionCopyright
Females pink, of typical mealybug appearance, often covered with white 'meal', up to 7 mm long.
TitleAdults on stem
CaptionFemales pink, of typical mealybug appearance, often covered with white 'meal', up to 7 mm long.
CopyrightPeter G. Allsopp
Females pink, of typical mealybug appearance, often covered with white 'meal', up to 7 mm long.
Adults on stemFemales pink, of typical mealybug appearance, often covered with white 'meal', up to 7 mm long.Peter G. Allsopp
S. sacchari adults parasitized by Anagyrus saccharicola.
TitleParasitized adult
CaptionS. sacchari adults parasitized by Anagyrus saccharicola.
CopyrightPeter G. Allsopp
S. sacchari adults parasitized by Anagyrus saccharicola.
Parasitized adultS. sacchari adults parasitized by Anagyrus saccharicola.Peter G. Allsopp
S. sacchari adults parasitized by Aspergillus flavus.
TitleParasitized adults
CaptionS. sacchari adults parasitized by Aspergillus flavus.
CopyrightPeter G. Allsopp
S. sacchari adults parasitized by Aspergillus flavus.
Parasitized adultsS. sacchari adults parasitized by Aspergillus flavus.Peter G. Allsopp

Identity

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Preferred Scientific Name

  • Saccharicoccus sacchari (Cockerell)

Preferred Common Name

  • grey sugarcane mealybug

Other Scientific Names

  • Dactylopius sacchari Ckll.
  • Dactylopius sacchari brasiliensis van Gorkum, 1913
  • Erium sacchari (Cockerell)
  • Phenacoccus sacchari
  • Pseudococcus sacchari Ckll.
  • Pseudococcus sacchari brasiliensis Gorkum
  • Pseudococcus saccharina
  • Trechocorys sacchari Cockerell
  • Trionymus praegrandis James, 1936
  • Trionymus sacchari Ckll.
  • Tryonymus sacchari Cockerell

International Common Names

  • English: grey mealybug of sugarcane; pink sugarcane mealy bug; sugarcane mealybug; sugarcane, mealybug; sugarcane, mealybug, pink
  • Spanish: chinche harinosa de la cana de azucar; chinche harinosa gris; chinche harinosa gris de la caña; chinche rosada de la cana de azucar; cochinilla blanca de la caña de azúcar
  • French: cochenille de la canne a sucre; cochenille grise de la canne à sucre

Local Common Names

  • Brazil: cochonilha rosada; cochonilha rosada da cana
  • Germany: Schmierlaus, Zuckerrohr-
  • Italy: Cocciniglia della Canna da zucchero
  • Japan: Kansyo-konakaigaramusi
  • South Africa: suikerriet-wolluis

EPPO code

  • PSECSA (Pseudococcus sacchari)

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta
  •                     Order: Hemiptera
  •                         Suborder: Sternorrhyncha
  •                             Unknown: Coccoidea
  •                                 Family: Pseudococcidae
  •                                     Genus: Saccharicoccus
  •                                         Species: Saccharicoccus sacchari

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Continent/Country/RegionDistributionLast ReportedOriginFirst ReportedInvasiveReferenceNotes

Asia

BangladeshPresentBen-Dov, 1994
CambodiaPresentCIE, 1977
ChinaPresentWilliams, 1970
-GuangdongPresentCIE, 1977
-Hong KongPresentNHM London UK 1996
-TibetPresentBen-Dov, 1994
Cocos IslandsPresentBellis et al., 2004
IndiaPresentPresent based on regional distribution.
-Andhra PradeshPresentCIE, 1977
-BiharPresentCIE, 1977
-Indian PunjabPresentCIE, 1977
-KarnatakaPresentCIE, 1977
-Madhya PradeshPresentCIE, 1977
-MaharashtraPresentCIE, 1977
-OdishaPresentCIE, 1977
-Tamil NaduPresentCIE, 1977
-Uttar PradeshPresentCIE, 1977
-West BengalPresentCIE, 1977
IndonesiaPresentWaterhouse, 1993
-Irian JayaPresentWilliams, 1970
-JavaPresentWilliams, 1970
IsraelPresentWilliams, 1970
JapanPresentWilliams, 1970
-Ryukyu ArchipelagoPresentWilliams, 1970
LaosPresentCIE, 1977
MalaysiaPresentWaterhouse, 1993
-Peninsular MalaysiaPresentWilliams, 1970
-SabahPresentCIE, 1977
-SarawakPresentWilliams, 1970
NepalPresentPradhan, 1981
OmanPresentNHM London UK 1996
PakistanPresentWilliams, 1970
PhilippinesPresentWilliams, 1970
SingaporePresentCIE, 1977
Sri LankaPresentWilliams, 1970
TaiwanPresentWilliams, 1970
ThailandPresentNapompeth, 1978
VietnamPresentCIE, 1977

Africa

AngolaPresentBen-Dov, 1994
EgyptPresentWilliams, 1970
EthiopiaPresentNHM London UK 1996
GhanaPresentWilliams, 1970
KenyaPresentWilliams, 1970
MadagascarPresentWilliams, 1970
MalawiPresentWilliams, 1970
MauritiusPresentWilliams, 1970
MozambiquePresentWilliams, 1970
NamibiaPresentNHM London UK 1996
RéunionPresentWilliams, 1970
SomaliaPresentWilliams, 1970
South AfricaPresentWilliams, 1970
SudanPresentNHM London UK 1996
SwazilandPresentCIE, 1977
TanzaniaPresentWilliams, 1970
UgandaPresentWilliams, 1970
ZimbabwePresentWilliams, 1970

North America

MexicoPresentWilliams, 1970
USAPresentPresent based on regional distribution.
-FloridaPresentHall et al., 2005
-HawaiiPresentWilliams, 1970
-TexasPresentMeagher et al., 1993

Central America and Caribbean

Antigua and BarbudaPresentWilliams, 1970
BahamasPresentWilliams, 1970
BarbadosPresentWilliams, 1970
British Virgin IslandsPresentWilliams, 1970
Cayman IslandsPresentWilliams, 1970
Costa RicaPresentWilliams, 1970
CubaPresentWilliams, 1970
Dominican RepublicPresentWilliams, 1970
El SalvadorPresentWilliams, 1970
GrenadaPresentWilliams, 1970
GuadeloupePresentWilliams, 1970
GuatemalaPresentWilliams, 1970
HondurasPresentWilliams, 1970
JamaicaPresentWilliams, 1970
MartiniquePresentWilliams, 1970
MontserratPresentWilliams, 1970
NicaraguaPresentWilliams, 1970
PanamaPresentWilliams, 1970
Puerto RicoPresentWilliams, 1970
Saint Kitts and NevisPresentWilliams, 1970
Saint LuciaPresentWilliams, 1970
Saint Vincent and the GrenadinesPresentWilliams, 1970
Trinidad and TobagoPresentWilliams, 1970
United States Virgin IslandsPresentCIE, 1977

South America

ArgentinaPresentWilliams, 1970
BoliviaPresentWilliams, 1970
BrazilPresentPresent based on regional distribution.
-AlagoasPresentCIE, 1977
-AmazonasPresentCIE, 1977
-Mato GrossoPresentCIE, 1977
-Minas GeraisPresentCIE, 1977
-ParaPresentCIE, 1977
-ParaibaPresentCIE, 1977
-PernambucoPresentCIE, 1977
-Rio de JaneiroPresentCIE, 1977
-Santa CatarinaPresentCIE, 1977
-Sao PauloPresentCIE, 1977
ColombiaPresentWilliams, 1970
EcuadorPresentWilliams, 1970
GuyanaPresentWilliams, 1970
ParaguayPresentWilliams, 1970
PeruPresentWilliams, 1970
SurinamePresentWilliams, 1970
UruguayPresentBen-Dov, 1994
VenezuelaPresentWilliams, 1970

Europe

PortugalPresentPresent based on regional distribution.
-MadeiraPresentWilliams, 1970
SpainPresentWilliams, 1970

Oceania

American SamoaPresentWilliams and Watson, 1988
AustraliaPresentWilliams, 1970
-New South WalesPresentAllsopp et al., 1993a
-QueenslandPresentAllsopp et al., 1993a
FijiPresentWilliams, 1970
French PolynesiaPresentWilliams, 1970
GuamPresentCIE, 1977
Micronesia, Federated states ofPresentBen-Dov, 1994
New CaledoniaPresentWilliams, 1970
Northern Mariana IslandsPresentWilliams, 1970
PalauPresentCIE, 1977
Papua New GuineaPresentWilliams, 1970
SamoaPresentWilliams, 1970
Solomon IslandsPresentBen-Dov, 1994
TongaPresentCIE, 1977
VanuatuPresentWilliams and Watson, 1988

Growth Stages

Top of page Pre-emergence, Vegetative growing stage

List of Symptoms/Signs

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SignLife StagesType
Leaves / abnormal colours
Leaves / honeydew or sooty mould
Leaves / honeydew or sooty mould
Roots / external feeding
Stems / external feeding

Natural enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Abdastartus atrus Predator
Anagyrus diversicornis Parasite Gujarat
Anagyrus saccharicola Parasite Adults/Nymphs Australia; Barbados; Bolivia; Fiji; Hawaii; Peru; St Kitts Nevis sugarcane
Anagyrus swezeyi Parasite
Angyrus saccharicola
Aspergillus flavus Antagonist Bolivia sugarcane
Aspergillus parasiticus Parasite Adults/Nymphs
Astymachus japonicus Parasite Adults/Nymphs
Brachiacantha Predator
Brachiacantha sp. Predator Adults/Nymphs
Carpophilus marginellus Predator
Chartocerus fimbriae Parasite
Chartocerus sp. Hyperparasite
Chilocorus nigrita Predator Adults/Nymphs
Chilocorus nigritus Predator Adults/Nymphs
Cryptolaemus montrouzieri Predator
Domomyza perspicax Predator Adults/Nymphs
Entomophthora pseudococci Pathogen
Exochomus flavipes Predator Adults/Nymphs
Exochomus flavipes Predator Adults/Nymphs
Exochomus melanocephalus Predator Egypt sugarcane
Hyperaspis limbatus Predator Hawaii sugarcane
Hyperaspis maindroni Predator
Hyperaspis notata Predator Adults/Nymphs
Hyperaspis trilineata Predator Adults/Nymphs Hawaii; St Kitts Nevis sugarcane
Millardia meltada Predator
Oligota barbadorum Predator
Oplobates woodwardi Predator
Pseudaphycus mundus Parasite Adults/Nymphs
Rhopus mymaricoides Parasite
Rodolia amabilis Predator
Scymnus apiciflavus Predator Hawaii sugarcane
Scymnus pallidivestris Predator Adults/Nymphs

References

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Alam MM, 1972. The establishment of Anagyrus saccharicola Timb.(Hymenoptera: Encyrtidae) in Barbados, West Indies, against the sugarcane mealybug, Saccharicoccus sacchari (Ckll.) (Hemiptera:Coccidae). Entomophage, 17(4):357-363

Allsopp PG, 1991. Binomial sequential sampling of adult Saccharicoccus sacchari on sugarcane. Entomologia Experimentalis et Applicata, 60(3):213-218

Allsopp PG; Chandler KJ; Samson PR; Story PG, 1993a. Pests of Australian Sugarcane. Brisbane, Australia: Bureau of Sugar Experiment Stations.

Allsopp PG; Sullivan GT; Haysom MBC; Morgan TA, 1993. Relationship of edaphic factors, location and harvest date to population levels of Saccharicoccus sacchari (Hemiptera: Pseudococcidae) on sugarcane. Environmental Entomology, 22(6):1278-1284

Ammar IMA; El-Nabawi A; Eisa A, 1986. Effect of two plant growth regulators (PGRs) on the mealybug and stem borers infestation on the sugar cane, in relation to the plant response and the yield. Annals of Agricultural Science, Moshtohor, 24(3):1679-1687

Anwar T; Jabbar A; Khalique F; Tahir S; Shakeel MA, 1992. Plants with insecticidal activities against four major insect pests in Pakistan. Tropical Pest Management, 38(4):431-437

APPPC, 1987. Insect pests of economic significance affecting major crops of the countries in Asia and the Pacific region. Technical Document No. 135. Bangkok, Thailand: Regional Office for Asia and the Pacific region (RAPA).

Ashbolt NJ; Inkerman PA, 1990. Acetic acid bacterial biota of the pink sugar cane mealybug, Saccharicoccus sacchari, and its environs. Applied and Environmental Microbiology, 56(375):707-712; 45 ref.

Atiqui MUA; Murad H, 1990. Morphology and ultrastructure of the ovary of Saccharicoccus sacchari Ckll. (Homoptera: Pseudococcidae) [a pest of sugarcane in India]. Zoologische Jahrbucher, Abteilung fur Anatomie und Ontogenie der Tiere, 120(3):197-209

Atiqui MUA; Murad H, 1992. Assessment of loss in sucrose content of sugarcane due to sugarcane mealy bug, Saccharicoccus sacchari Ckll. Journal of Insect Science, 5(2):196-197

Barro PJde, 1990. Natural enemies and other species associated with Saccharicoccus sacchari (Cockerell) (Hemiptera: Pseudococcidae) in the Bundaberg area, southeast Queensland. Journal of the Australian Entomological Society, 29(2):87-88; 6 ref.

Barro PJde, 1991. Sampling strategies for above and below ground populations of Saccharicoccus sacchari (Cockerell) (Hemiptera: Pseudococcidae) on sugarcane. Journal of the Australian Entomological Society, 30(1):19-20

Bartlett BR, 1977. Pseudococcidae. In: Clausen CP, ed. Introduced Parasites and Predators of Arthropod Pests and Weeds: A World Review. Agriculture Handbook, US Department of Agriculture No. 480, 137-170.

Beardsley JW, 1959. Observations on sugar cane mealybugs in Hawaii. Proceedings of the International Society of Sugar Cane Technologists, 10:954-961.

Beardsley JW, 1960. A preliminary study of the males of some Hawaiian mealybugs (Homoptera: Pseudococcidae). Proceedings of the Hawaiian Entomological Society, 17:199-243.

Bellis GA; Donaldson JF; Carver M; Hancock DL; Fletcher MJ, 2004. Records of insect pests on Christmas Island and the Cocos (Keeling) Islands, Indian Ocean. Australian Entomologist, 31(3):93-102.

Ben-Dov Y, 1994. A systematic catalogue of the mealybugs of the world (Insecta: Homoptera: Coccoidea: Pseudococcidae and Putoidae) with data on geographical distribution, host plants, biology and economic importance. Andover, UK; Intercept Limited, 686 pp.

Box HE, 1953. List of sugar-cane Insects. London, UK: Commonwealth Institute of Entomology.

Bynum EK, 1937. Pseudococcobius terryi Fullaway: a Hawaiian parasite of grey sugarcane mealybug in the United States. Journal of Economic Entomology, 30:756-761.

Carver M; Inkerman PA; Ashbolt NJ, 1987. Anagyrus saccharicola Timberlake (Hymenoptera: Encyrtidae) and other biota associated with Saccharicoccus sacchari (Cockerell) (Homoptera: Pseudococcidae) in Australia. Journal of the Australian Entomological Society, 26(4):367-368; 5 ref.

CIE, 1977. Distribution Maps of Plant Pests, No. 102. Wallingford, UK: CAB International.

Clancy DW; Pollard HN, 1947. Further experiments in the parasitization of mealybugs. Journal of Economic Entomology, 40:578-579.

Cock MJW (ed.), 1985. A review of biological control of pests in the Commonwealth Caribbean and Bermuda up to 1982. Farnham Royal, United Kingdom; Commonwealth Agricultural Bureaux, xii + 218 pp.

Coles LW, 1964. Collection of the predacious lady beetle, Hyperaspis trilineata, in Barbados, and shipment to Hawaii. Journal of Economic Entomology, 57:768-769.

Compere H; Subba Rao BR; Kaur RB, 1959. Two species of Encyrtidae parasitic in the pink mealybug of sugar cane in India. Proceedings of the National Institute of Science of India (B), 26:45-50.

Cooper DJ; Zhang QY, 1992. Virus-like particles in the pink sugarcane mealybug, Saccharicoccus sacchari. Journal of Invertebrate Pathology, 60(2):206-207

Dick J, 1969. The mealybugs of sugar cane. In: Williams JR, Metcalfe JR, Mungomery RW, Mathes R, eds. Pests of Sugar Cane. Amsterdam, Netherlands: Elsevier, 343-365.

Drummond J; Barro PJde; Pinnock DE, 1991. Field and laboratory studies on the fungus Aspergillus parasiticus, a pathogen of the pink sugar cane mealybug Saccharicoccus sacchari.. Biological Control, 1(4):288-292; 12 ref.

Drummond J; Pinnock DE, 1990. Aflatoxin production by entomopathogenic isolates of Aspergillus parasiticus and Aspergillus flavus.. Journal of Invertebrate Pathology, 55(3):332-336; 15 ref.

Duhra MS; Singh DP, 1986. Relative toxicity of some insecticides to the sugar cane mealy bug. Sugarcane, No. 6:10-11

Dymond; GC, 1929. Mealybugs. Their effect on cane culture and manufacture in Natal. Proceedings of the South African Sugarcane Technologists Association, 3:51-53.

Gahan AB, 1946. Eight new species of chalcid-flies of the genus Pseudaphycus with a key to species. Proceedings of the US National Museum, 96:311-327.

Hafez M; Salama HS, 1967. Polymorphism in the male of the mealy bug Saccharicoccus sacchari. Proceedings of the Royal Entomological Society of London A, 42:25-29.

Hafez M; Salama HS, 1969a. Ecological studies on the sugar cane mealybug Saccharicoccus sacchari Ckll., in Egypt (Homoptera: Coccoidea). Bulletin de la Societe Entomologique d'Egypte, 53:21-39.

Hafez M; Salama HS, 1969b. Temperature and humidity reactions of the sugar cane mealybug, Saccharicoccus sacchari Ckll. (Homoptera: Coccoidea). Bulletin de la Societe Entomologique d'Egypte, 53:41-62.

Hafez M; Salama HS, 1969c. Biological studies on the sugar cane mealy bug, Saccharicoccus sacchari Ckll., in Egypt. Bulletin de la Societe Entomologique d'Egypte, 53:499-516.

Hall DG, 1988. Insects and mites associated with sugarcane in Florida. Florida Entomologist, 71(2):138-150

Hall DG; Konstantinov AS; Hodges GS; Sosa O; Welbourn C; Westcott RL, 2005. Insects and mites new to Florida sugarcane. Journal - American Society of Sugar Cane Technologists, 25:143-156. http://www.assct.org/journal/JASSCT%20PDF%20Files/vol25/A05-07%20Hall%20final.pdf

Hall WJ, 1922. The outbreaks of Pseudococcus sacchari Ckll. on the sugar cane of Egypt. Bulletin Minist. Agric. Egypt Tech. Scient. Serv., 26:1-16.

Lockhart BEL; Autrey LJC; Comstock JC, 1992. Partial purification and serology of sugarcane mild mosaic virus, a mealybug-transmitted closterolike virus. Phytopathology, 82(6):691-695

Mahmood R; Mohyuddin AI; Murtaza M, 1982. Pseudococcids attacking sugar-cane in Pakistan and their natural enemies. Proceedings of the Annual Convention of the Pakistan Society of Sugar Technologists, 193-205.

Mathes R; Ingram JW; Questel DD; Thames WH; Dugas AL, 1953. Current status of sugarcane-insect investigations in the United States. Proceedings of the International Society of Sugar Cane Technologists, 7:560-567.

Meagher RL Jr; Wilson SW; Blocker HD; Eckel RVW; Pfannenstiel RS, 1993. Homoptera associated with sugarcane fields in Texas. Florida Entomologist, 76(3):508-514

Mesbah HA; Fahmy IS; El-Deeb AS; Gaber AA; Nour AH, 1976. The susceptibility of ten sugar cane varieties to infestation with borers and mealybugs at Alexandria, Egypt. Bulletin of the Entomological Society of Egypt, No. 60:403-411

Myo Chit, 1988. Comparison of biological attributes of Anagyrus saccharicola Timb. (Hymenoptera: Encyrtidae) and Pseudaphycus mundus Gahan (Hymenoptera: Encyrtidae), parasitoids of pink sugarcane mealybug, Saccharicoccus sacchari Ckll. (Homoptera: Pseudococcidae). MS dissertation, Texas A&M University, College Station, Texas, USA.

Napompeth B(Project leader), 1978. Sugarcane insect pest management. Report, Kasetsart University, Thailand, 66-67

Nigam H, 1983. Complex of biotic agents associated with pink mealy bug of sugarcane, Saccharicoccus sacchari Gkll. in Tarai region of Uttar Pradesh. Indian Journal of Entomology, 45(3):321

Noyes JS; Hayat M, 1994. Oriental mealybug parasitoids of the Anagyrini (Hymenoptera: Encyrtidae). Wallingford, UK; CAB International, viii + 554 pp.

Pradhan SB, 1981. Rice mealybug and its alternate host plants. International Rice Research Newsletter, 6(4):11-12

Rae DJ, 1993. A method for discrimination between instars of Saccharicoccus sacchari (Cockerell) (Hemiptera: Pseudococcidae). Journal of the Australian Entomological Society, 32(3):249-252

Rae DJ; De'ath G, 1991. Influence of constant temperature on development, survival and fecundity of the sugarcane mealybug, Saccharicoccus sacchari (Cockerell) (Hemiptera: Pseudococcidae). Australian Journal of Zoology, 39:105-122.

Rae DJ; Jones RE, 1992. Influence of host nitrogen levels on development, survival, size and population dynamics of sugarcane mealybug, Saccharicoccus sacchari (Cockerell) (Hemiptera: Pseudococcidae). Australian Journal of Zoology, 40(3):327-342

Rajendra A, 1974. The biology and control of Saccharicoccus sacchari Ckll. (Hom: Pseudococcidae) the pink mealy bug of sugar cane in Sri Lanka. Ceylon Journal of Science, Biological Sciences, 11(1):23-28

Rezende JAM; Rossetto CJ; Germeck EB, 1979. Incidence of Saccharicoccus sacchari (Cockerell, 1895) (Homoptera, Coccoidea, Pseudococcidae) in varieties of sugar-cane. Bragantia, 38:XXXVII-XL

Rezende JAM; Rossetto CJ; Germeck EB, 1979. Infestation of Saccharicoccus sacchari (Cockerell, 1895) in different varieties of Saccharum spp. Entomology Newsletter, International Society of Sugarcane Technologists, No. 6:18

Uichanco LB; Villaneuva FE, 1932. Biology of the pink mealy bug of sugar cane, Trionymus sacchari (Cockerell), in the Philippines. Philippines Agriculturalist, 21:205-276.

van Dine DL, 1913. The insects affecting sugar cane in Puerto Rico. Journal of Economic Entomology, 6:251-255.

Waterhouse DF, 1993. The Major Arthropod Pests and Weeds of Agriculture in Southeast Asia. ACIAR Monograph No. 21. Canberra, Australia: Australian Centre for International Agricultural Research, 141 pp.

Williams DJ, 1962. The British Pseudococcidae (Homoptera: Coccoidea). Bulletin of the British Museum (Natural History) (Entomology), 8:385-430.

Williams DJ, 1970. The mealy bugs (Homoptera: Coccoidea: Psuedococcidae) of sugarcane, rice and sorghum. Bulletin of Entomological Research, 60:109-188.

Williams DJ, 1985. Australian mealybugs. London, UK; British Museum (Natural History), 431 pp.

Williams DJ; Granara de Willink MC, 1992. Mealybugs of Central and South America. Wallingford, UK: CAB International.

Williams DJ; Watson GW, 1988. Scale insects of the tropical South Pacific region. Part 2. Mealybugs (Pseudococcidae). Wallingford, Oxon, UK; CAB International, 260 pp.

Yadava RL, 1966. Morphology of the male pink sugarcane mealybug, Saccharicoccus sacchari Cockerell (Pseudococcidae: Homoptera). I. Macropterous form. Agra University Journal of Research, 15:71-129.

Yadava RL, 1967a. Morphology of the male pink sugarcane mealybug, Saccharicoccus sacchari Ckll (Pseudococcidae: Homoptera). II. Apterous form. Agra University Journal of Research, 16:101-133.

Yadava RL, 1967b. Morphology of the male pink sugarcane mealybug, Saccharicoccus sacchari Cockerell (Pseudococcidae: Homoptera). III. Brachypterous form. Agra University Journal of Research, 16:141-172.

Distribution Maps

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