Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide

Datasheet

Morus alba
(mora)

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Datasheet

Morus alba (mora)

Summary

  • Last modified
  • 22 November 2019
  • Datasheet Type(s)
  • Invasive Species
  • Host Plant
  • Preferred Scientific Name
  • Morus alba
  • Preferred Common Name
  • mora
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Plantae
  •     Phylum: Spermatophyta
  •       Subphylum: Angiospermae
  •         Class: Dicotyledonae
  • Summary of Invasiveness
  • M. alba is a multipurpose tree widely planted in tropical, subtropical and mild temperate regions of the world for fodder and silkworm rearing, and for fruit and timber production (

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Pictures

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PictureTitleCaptionCopyright
Morus alba (mora), habit, on farmland.
TitleTree habit
CaptionMorus alba (mora), habit, on farmland.
Copyright©K.M. Siddiqui
Morus alba (mora), habit, on farmland.
Tree habitMorus alba (mora), habit, on farmland.©K.M. Siddiqui
Morus alba (mora); sapling.
TitleSapling
CaptionMorus alba (mora); sapling.
Copyright©K.M. Siddiqui
Morus alba (mora); sapling.
SaplingMorus alba (mora); sapling.©K.M. Siddiqui
Morus alba (mora); foliage and fruit.
TitleFoliage and fruit
CaptionMorus alba (mora); foliage and fruit.
Copyright©K.M. Siddiqui
Morus alba (mora); foliage and fruit.
Foliage and fruitMorus alba (mora); foliage and fruit.©K.M. Siddiqui
Morus alba (mora); nursery seedlings.
TitleSeedlings
CaptionMorus alba (mora); nursery seedlings.
Copyright©K.M. Siddiqui
Morus alba (mora); nursery seedlings.
SeedlingsMorus alba (mora); nursery seedlings.©K.M. Siddiqui

Identity

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Preferred Scientific Name

  • Morus alba L.

Preferred Common Name

  • mora

Other Scientific Names

  • Morus alba f. tatarica Ser.
  • Morus alba var. constantinopolitana Loudon
  • Morus alba var. multicaulis (Perr.) Loudon
  • Morus atropurpurea Roxb.
  • Morus indica L.
  • Morus multicaulis Perr.

International Common Names

  • English: mulberry; white mulberry; white mulberry tree
  • Spanish: morera
  • French: murier; Murier blanc; mûrier blanc
  • Arabic: tuth

Local Common Names

  • Brazil: amora-branca; amoreira-branca; amoreira-preta
  • Germany: Weisser Maulbeerbaum
  • India: ambat; chinni; hipnerle; kamblichedi; musukette; pippalipandu chettu; reshme chattu; reshms chattu; shahtut; shehtun; shetur; siah tut; tula; tut; tuticoli; tuto; tutri
  • Italy: Gelso bianco; moral blanco; morera blanco
  • Netherlands: moerbei, witte
  • Pakistan: chinni; shahtut; tut; tutri

EPPO code

  • MORAL (Morus alba)

Trade name

  • mulberry
  • tut
  • white mulberry

Summary of Invasiveness

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M. alba is a multipurpose tree widely planted in tropical, subtropical and mild temperate regions of the world for fodder and silkworm rearing, and for fruit and timber production (Gupta, 1993). Introduced M. alba has been found to be associated with invasion events in Oregon and Virgina (USA) and South Africa (Haysom and Murphy, 2003). It is also recorded as invasive in Brazil (Instituto Horus, 2011).

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Plantae
  •         Phylum: Spermatophyta
  •             Subphylum: Angiospermae
  •                 Class: Dicotyledonae
  •                     Order: Urticales
  •                         Family: Moraceae
  •                             Genus: Morus
  •                                 Species: Morus alba

Notes on Taxonomy and Nomenclature

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The genus Morus (family Moraceae) contains about 10 species (Parker, 1956). Many cultivars have been developed over the long period of time that Morus spp. have been cultivated for leaf and fruit production (Gerasopoulos and Stavroulakis, 1997). For example, there are about 700 cultivars in Japan that are used for rearing silkworms. There has been confusion regarding the characteristics of M. alba at various times (Watt, 1972).

Description

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A small to medium tree growing up to 15 m, M. alba has a short trunk, and a rounded crown with a dense canopy of spreading branches. Leaves are alternate, simple, 6-18 cm long, 5-13 cm wide, broadly ovate, dentate or lobed with 3 prominent veins running from the rounded or obliquely cordate base. Somewhat polymorphic, leaves are shiny green on the adaxial surface, paler and slightly hairy underneath. Bark is light brown to grey, smooth but may be furrowed. Dioecious, male inflorescences are small with 4 stamens, filaments inflexed in bud, green and borne on long catkins. Female flowers are inconspicuous, perianth with 4 free or almost free segments, aggregated in short spikes. Fruit is an ovoid or cylindrical syncarp composed of achenes, pendunculate, red when immature, blackish-purple, purple or greenish-white when mature, 1-2.5 cm long (Global Invasive Species Database, 2016; Tutin, 1964).

Two botanical varieties of M. alba are recognized (Zhou and Gilbert 2003): varieties ‘alba’ and ‘multicaulis’. The leaf blade of ‘alba’ is 5-15 cm, fruits are ovoid to ellipsoid, blackish purple when mature. The leaf blade of var. ‘multicaulis’ is <30 cm with cylindric fruits which are greenish-white to purple when mature.

Distribution

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M. alba is native to central and eastern China, and is widely cultivated and naturalized elsewhere across the world, largely due to sericulture. In some areas M. alba is invasive, hybridizing with native species M. rubra in North America and outcompeting other native species in South Africa.

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 17 Dec 2021
Continent/Country/Region Distribution Last Reported Origin First Reported Invasive Planted Reference Notes

Africa

EgyptPresentPlanted
MauritaniaPresentPlanted
MauritiusPresentPlanted
RéunionPresentIntroduced1775
South AfricaPresentIntroducedInvasivePlanted
TanzaniaPresentPresent based on regional distribution.
-Zanzibar IslandPresentPlanted
TunisiaPresentPlanted
ZambiaPresentPlanted

Asia

AfghanistanPresentPlanted
BhutanPresentIntroduced1965
ChinaPresentPlanted
-BeijingPresent
-GuangdongPresent
-GuangxiPresent
-HenanPresent
-JiangsuPresent
-JiangxiPresent
-ShandongPresent
-SichuanPresent
-TibetPresentPlanted
-ZhejiangPresent
IndiaPresentPlanted
-Jammu and KashmirPresent
-KarnatakaPresent
-Tamil NaduPresent
-Uttar PradeshPresent
IranPresentPlanted
IraqPresentPlanted
IsraelPresentPlanted
JapanPresentPlanted
JordanPresentPlanted
KazakhstanPresentPlanted
KyrgyzstanPresentPlanted
North KoreaPresentPlanted
PakistanPresentPlanted
South KoreaPresentPlanted
TaiwanPresentPlanted
TajikistanPresentPlanted
TurkeyPresentPlanted
TurkmenistanPresentPlanted
UzbekistanPresentPlanted

Europe

AlbaniaPresent
AustriaPresentIntroducedFirst reported: <1830
BelgiumPresentIntroducedFirst reported: <1850
BulgariaPresent
CroatiaPresent
CyprusPresentPlanted
Federal Republic of YugoslaviaPresentPlanted
Union of Soviet Socialist RepublicsPresent
France
-CorsicaPresentIntroduced1910
GreecePresentPlanted
HungaryPresentPlanted
ItalyPresentPlanted
-SicilyPresent
LithuaniaPresentIntroduced1990
PolandPresent
RomaniaPresent
RussiaPresentPlanted
SlovakiaPresentIntroducedFirst reported: 1720. First reported in wild: 1830
SloveniaPresentIntroduced1900
SpainPresent
United KingdomPresent

North America

Costa RicaPresent
United StatesPresentPlanted
-HawaiiPresentIntroduced1920
-OregonPresentIntroducedInvasive
-VirginiaPresentIntroducedInvasive

Oceania

AustraliaPresentIntroduced1843
-Lord Howe IslandPresentIntroduced1870

South America

ArgentinaPresentPlanted
BrazilPresentIntroducedInvasivePlanted

Habitat List

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CategorySub-CategoryHabitatPresenceStatus
Terrestrial

Biology and Ecology

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M. alba is a rapidly growing deciduous woody perennial with a deep root system. Mulberry trees are either dioecious or monoecious, and a tree sometimes will change from one sex to another. High temperatures, strong light and long days favour maleness in mulberries, with their opposites, as well as high humidity, favouring the production of female flowers. Flowers appear as cylindrical catkins along with the leaves axillary on the current season’s shoots and on spurs on older wood. Pollination is not usually a problem because mulberries are wind pollinated, and some plants will set fruit without any pollination whatsoever. In China, M. alba flowers in April to May with fruiting during May to August.

White mulberry is grown in cool temperate steppe to warm to tropical, very dry to moist, forest life zones and can be grown in annual temperatures ranging from 6-28°C (Duke, 1983). It is the most cold hardy of the mulberry species grown for their fruit, though cold hardiness varies from clone to clone. The clone ‘Tartarica’ is tolerant of temperatures down to -32°C. M. alba prefers moist, well drained loamy soils in a sunny position. However, it will grow in coarse, medium and fine soils, has some shade tolerance and is moderately tolerant of drought. Once established, it is tolerant of drought (Duke, 1983) and of saline conditions (Global Invasive Species Database, 2016). It is liable to wind damage, but in India it is cultivated up to 3300 m in altitude (Duke, 1983).

Latitude/Altitude Ranges

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Latitude North (°N)Latitude South (°S)Altitude Lower (m)Altitude Upper (m)
40 -10 250 3300

Air Temperature

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Parameter Lower limit Upper limit
Absolute minimum temperature (ºC) -28
Mean maximum temperature of hottest month (ºC) 21 37
Mean minimum temperature of coldest month (ºC) 9 19

Rainfall

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ParameterLower limitUpper limitDescription
Dry season duration24number of consecutive months with <40 mm rainfall
Mean annual rainfall12002000mm; lower/upper limits

Soil Tolerances

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Soil drainage

  • free

Soil reaction

  • neutral

Soil texture

  • heavy
  • light
  • medium

Uses

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Worldwide M. alba is mainly cultivated for sericulture. China has over 400,000 ha planted for their leaves as silkworm fodder. Leaves are also used as fodder for ruminant animals and have even been eaten as a vegetable by humans. The bark can be used for paper. Ripe fruits are eaten fresh, used in desserts and cooking in pies and tarts, and also for cordials and jams. In Asia, southern Europe and southern parts of the USA, M. alba has been used for windbreaks, in landscaping, and their low water requirements and resistance to pruning makes them suitable for use as a street tree. In India the wood is used for cabinet work and sporting goods (Sánchez, 2002).

Uses List

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Animal feed, fodder, forage

  • Fodder/animal feed
  • Invertebrate food for silkworms

Environmental

  • Agroforestry
  • Boundary, barrier or support
  • Erosion control or dune stabilization
  • Revegetation
  • Shade and shelter
  • Windbreak

Fuels

  • Charcoal

General

  • Ornamental

Human food and beverage

  • Honey/honey flora

Materials

  • Carved material
  • Fibre
  • Gum/resin
  • Wood/timber

Medicinal, pharmaceutical

  • Source of medicine/pharmaceutical
  • Traditional/folklore

Wood Products

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Charcoal

Containers

  • Boxes
  • Cases
  • Cooperage
  • Crates

Pulp

  • Short-fibre pulp

Sawn or hewn building timbers

  • Carpentry/joinery (exterior/interior)
  • For light construction

Wood-based materials

  • Plywood

Woodware

  • Industrial and domestic woodware
  • Musical instruments
  • Sports equipment
  • Tool handles
  • Turnery

References

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Ahmad I, Park KE, 1989. Mulberry Cultivation Techniques. Peshawar, Pakistan: Pakistan Forest Institute.

Benavides J, 1995. Management and utilization of mulberry (Morus alba) for forage in Central America. Agroforestería en las Américas, 2(7):27-30; 7 ref.

Berdyklycev K, 1968. The importance of woody species for drainage in the conditions of N. Turkmenia. [Meliorativnoe znacenie drevesnyh porod v uslovijah SevernojTurkmenii.] Izdatel'stvo 'Ylym', Ashkhabad. pp. 100. [158 refs.].

Bokhari AS, 1973. Revised working plan of Changa Manga plantation (Kasur Forest Division). Lahore, India: Government Printing.

California Rare Fruit Growers, 2016. Mulberry, Morus spp., Moraceae https://www.crfg.org/pubs/ff/mulberry.html

Carlowitz PGvon, 1991. Multipurpose trees and shrubs: sources of seeds and inoculants. Multipurpose trees and shrubs: sources of seeds and inoculants., vii + 328 pp.; 46 ref.

Champion HG, Seth SK, Khattak GM, 1965. Forest Types of Pakistan. Peshawar, Pakistan: Pakistan Forest Institute.

Coventry BO, 1925. Devastation of Changa Manga plantation by fungus attack. Indian Forester, 51(10):517-523.

Duke, J. A., 1983. Handbook of energy crops, West Lafayette, Indiana: Purdue University, Center for New Crops & Plants Products https://hort.purdue.edu/newcrop/duke_energy/Morus_alba.html

Gerasopoulos D, Stavroulakis G, 1997. Quality characteristics of four mulberry (Morus sp) cultivars in the area of Chania, Greece. Journal of the Science of Food and Agriculture, 73(2):261-264.

Ghosh RC, 1977. Handbook on afforestation techniques. New Delhi, India; Controller of Publications, Government of India. 411 pp.; 11 pl., 4 maps; 379 ref.

Guha SRD, Madan RN, 1962. Chemical pulps for writing and printing papers from Morus alba. Indian Forester, 88(1):64-66.

Gupta RK, 1993. Multipurpose trees for agroforestry and wasteland utilisation. Multipurpose trees for agroforestry and wasteland utilisation., xv + 562 pp.; [18 pp. of ref + refs in text].

Haysom K, Murphy S, 2003. The status of invasiveness of forest tree species outside their natural habitat: a global review and discussion paper. Rome, Italy: FAO. http://www.fao.org/DOCREP/006/J1583E/J1583E00.htm

Hocking D, ed. , 1993. Trees for drylands. New Delhi, India: Oxford and IBH.

Huang YuLan, Lan LiuYan, Hu ShengHu, Zhang ChaoLan, Feng JianLing, Tang Jian, Li XiaoFeng, 2015. Impact of organic fertilizers and iron-fertilizers on Morus alba L. chlorosis in paddy soils., Acta Agriculturae Zhejiangensis, 27(11):1984-1989 http://www.zjnyxb.cn/EN/abstract/abstract2543.shtml

Instituto Horus, 2011. Alien Invasive Species: Fact Sheets., Brazil. http://www.institutohorus.br/inf_fichas_eng

Invasive Species Specialist Group, 2016. Global Invasive Species Database http://www.iucngisd.org/gisd/

Janick, J., Paull, R. E., 2008. The encyclopedia of fruit & nuts.:xviii + 954 pp.

Kadambi K, Dabral SN, 1954. Air layering in forestry practice. Indian Forester, 80(11):721-724 + 1 plate.

Khan AH, Bokhari AS, 1970. Damage due to fungus diseases in Bhagat Reservoir Plantation, Lyallpur Forest Division. Pakistan Journal of Forestry, 20(3):293-311. [3 refs.].

Litz, R. E., 2004. Biotechnology of fruit and nut crops.:xxiv + 723 pp. http://www.cabi.org/cabebooks/ebook/20053001306

Lochynska, M., Oleszak, G., 2011. Multi-use of the white mulberry (Morus alba L.)., Ecological Questions, 15:91-95 http://www.degruyter.com/view/j/ecoq.2011.15.issue-1/v10090-011-0040-5/v10090-011-0040-5.xml?format=INT

Luna RK, 1996. Plantation trees. Delhi, India: International Book Distributors.

Manian K, Nagarajan M, Padmanabhan G, Annamalainathan K, 1995. Influence of bio-regulators on biomass production in mulberry (Morus alba). Madras Agricultural Journal, 82(2):85-87; 8 ref.

Naowaratwattana, W., De-Eknamkul, W., Mejia, E. G. de, 2010. Phenolic-containing organic extracts of mulberry (Morus alba L.) leaves inhibit HepG2 hepatoma cells through G2/M phase arrest, induction of apoptosis, and inhibition of topoisomerase IIa activity., Journal of Medicinal Food, 13(5):1045-1056 http://www.liebertonline.com/jmf

Narayanaswamy KC, Geethabai M, Raghuraman R, 1996. Mite pests of mulberry - a review. Indian Journal of Sericulture, 35(1):1-8.

Ninova D, 1970. Anatomical indications of fume resistance in certain woody plants. Gorskostop. Nauka, 7(4):9-18. [13 refs.].

Oku, T., Yamada, M., Nakamura, M., Sadamori, N., Nakamura, S., 2006. Inhibitory effects of extractives from leaves of Morus alba on human and rat small intestinal disaccharidase activity., British Journal of Nutrition, 95(5):933-938

Oz, A. T., Ulukanli, Z., 2014. The effects of calcium chloride and 1-methylcyclopropene (1-MCP) on the shelf life of mulberries (Morus alba L.)., Journal of Food Processing and Preservation, 38(3):1279-1288 http://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1745-4549

Park KwangJun, Park KJ, 1995. Characteristics of several tetraploid induced by colchicine treatment in mulberry seedlings (Morus alba L. selfing Ist generation of Kaeryangppong). RDA Journal of Agricultural Science, Farm Management, Agricultural Engineering, Sericulture, and Farm Products Utilization, 37(2):759-765; 25 ref.

Parker RN, 1956. A forest flora for the Punjab with Hazara and Delhi. Lahore, Pakistan: Government Printing Press.

Petkov Z, 1995. Effect of some growth regulators on the germination of white mulberry (Morus alba L.) seeds. Rasteniev"dni Nauki, 32(7-8):149-152; 10 ref.

Porto, A. J., Costa, C., 2011. Postharvest storage systems for mulberry branches (Morus spp.), Veterinária e Zootecnia , 18(1):135-146

Quraishi MA, Ahmad M, 1971. New fungal records on Morus alba Linn. Pakistan Journal of Forestry, 21(3):303-312. [8 ref.].

Ramanjulu S, Sudhakar C, 1997. Drought tolerance is partly related to amino acid cumulation and ammonia assimilation: a comparative study in two mulberry genotypes differing in drought sensitivity. Journal of Plant Physiology, 150(3):345-35.

Royal Horticultural Society, 2016. Morus alba 'Pendula', weeping white mulberry https://www.rhs.org.uk/Plants/46790/Morus-alba-Pendula/Details

Sánchez, M. D., 2002. World distribution and utilization of mulberry and its potential for animal feeding., FAO Animal Production and Health Paper:1-9

Santhi P, Ponnuswamy K, Radha NV, 1994. Effect of foliar spray of micronutrients on different growth stages of mulberry (Morus alba Linn.,). Madras Agricultural Journal, 81(11):622-624; 5 ref.

Satiya Sharma, Mira Madan, 1994. Potential of mulberry (Morus alba) biomass., Journal of Scientific & Industrial Research, 53(9):710-714

Sheikh MI, 1993. Trees of Pakistan. GOP-USAID Forestry Planning and Development Project. Islamabad, Pakistan: Pictorial Printers (Pvt.) Ltd.

Shen Yan, Wang DongLan, Liu XianJin, Wu ChangFu, Sun Xing, Ma YaoYao, 2012. Residue characteristics of propargite in mulberry (Morus alba L.) and soil., Journal of Nanjing Forestry University (Natural Sciences Edition), 36(4):93-97 http://njlydxxb.periodicals.net.cn/default.html

Siddiqui KM, 1993. Irrigated Forestry in Pakistan. (A state-of-art-review). Peshawar, Pakistan: Pakistan Forest Institute.

Siddiqui KM, Ayaz M, Mahmood I, 1996. Properties and uses of Pakistan timbers. Peshawar, Pakistan: Forest Products Research Division, Pakistan Forest Institute.

Singab, A. N. B., El-Beshbishy, H. A., Yonekawa, M., Nomura, T., Fukai, T., 2005. Hypoglycemic effect of Egyptian Morus alba root bark extract: effect on diabetes and lipid peroxidation of streptozotocin-induced diabetic rats., Journal of Ethnopharmacology, 100(3):333-338

Singh SP, 1989. Wasteland development. New Delhi, India: Agricole.

Sosef MSM, Hong LT, Prawirohatmodjo S, eds, 1998. Plant resources of southeast Asia. Timber trees: lesser-known timbers. Leiden, The Netherlands: Backhuys Publishers, 5(3).

Streets RJ, 1962. Exotic forest trees in the British Commonwealth. Oxford, UK: Clarendon Press.

Tutin, T. G., 1964. Morus L. In: Tutin TG, Heywood VH, Burges NA, Valentine DH, Walters SM, Webb DA, eds. Flora Europea volume 1, Lycopodiaceae to Plantanaceae

Zhou, Z. K., Gilbert, M. G., 2003. Moraceae In: Wu ZY, Raven PH, Hong DY, eds. Flora of China. Volume 5 (Ulmaceae through Basellaceae:21-73 http://flora.huh.harvard.edu/china/mss/volume05/Moraceae.pdf

Distribution References

Abdel-Salam A M M, Rezk A A, Dawoud R A, 2019. Biochemical, serological, molecular and natural host studies on Tomato Chlorosis Virus in Egypt. Pakistan Journal of Biological Sciences. 22 (2), 83-94. https://scialert.net/fulltext/?doi=pjbs.2019.83.94&org=11

Abdul-Rassoul M S, 2014. Host plants of the mealybug Nipaecoccus viridis (Newstead, 1894) (Homoptera, Pseudococcidae) in Iraq with detection of new hosts. Advances in Bio Research. 5 (4), 3-6. http://soeagra.com/abr/abrdec2014/2.pdf

Abdul-Rassoul M S, 2015. Host plants of the mealybug Nipaecoccus viridis (Newstead, 1894) (Homoptera, pseudococcidae) in Iraq with detection of new hosts. Advances in Bio Research. 6 (2), 23-26. http://soeagra.com/abr/abrmarch2015/4.pdf

Alishiri A, Rakhshandehroo F, Shams-Bakhsh M, Jouzani G R S, 2016. Incidence and distribution of fig badnavirus 1 and mulberry badnavirus 1 on mulberry trees in Iran. Journal of Plant Pathology. 98 (2), 341-345. http://www.sipav.org/main/jpp/index.php/jpp/article/view/3509

Allahverdi T, Rahimian H, Ravanlou A, 2016. First report of bacterial canker in mulberry caused by Citrobacter freundii in Iran. Plant Disease. 100 (8), 1774-1775. http://apsjournals.apsnet.org/loi/pdis DOI:10.1094/PDIS-01-16-0020-PDN

Bjeliš M, Buljubašić I, Popović L, Milek T M, 2015. Spread of the spotted wing drosophila - Drosophila suzukii (Diptera, Drosophilidae) and new distribution records in Dalmatia region of Croatia. Bulletin OEPP/EPPO Bulletin. 45 (2), 214-217. http://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1365-2338

CABI Data Mining, Undated. CAB Abstracts Data Mining.,

CABI, Undated. CABI Compendium: Status inferred from regional distribution. Wallingford, UK: CABI

CABI, Undated a. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI

Castillo P, Vito M di, Vovlas N, Jiménez-Díaz R M, 2001. Host-parasite relationships in root-knot disease of white mulberry. Plant Disease. 85 (3), 277-281. DOI:10.1094/PDIS.2001.85.3.277

Chen J, Xiang T T, Liu X Y, Wang W H, Zhang B L, Liu J, Zhou W, Wan Y J, Chen G, Zhu H S, 2018. First report of Nigrospora sphaerica causing shot hole disease on mulberry in China. Plant Disease. 102 (1), 245-246. DOI:10.1094/PDIS-06-17-0914-PDN

Chen S C, Zhao H J, Wang M M, Jiang Q Y, Wang Z H, Liu A R, Ahammed G J, 2017. First report of stem rot caused by Coniothyrium fuckelii on Morus alba in China. Plant Disease. 101 (2), 387-388. DOI:10.1094/PDIS-07-16-1044-PDN

Chen S C, Zhao H J, Wang M M, Ren J J, Wang Z H, Li J D, Liu A R, Ahammed G J, 2017a. First report of root rot caused by Fusarium incarnatum on Morus alba in China. Plant Disease. 101 (7), 1320. DOI:10.1094/PDIS-02-17-0161-PDN

Haysom K, Murphy S, 2003. The status of invasiveness of forest tree species outside their natural habitat: a global review and discussion paper. In: The status of invasiveness of forest tree species outside their natural habitat: a global review and discussion paper, Rome, Italy: FAO. http://www.fao.org/DOCREP/006/J1583E/J1583E00.htm#TopOfPage

Șİșman S, Ülgentürk S, 2010. Scale insects species (Hemiptera: Coccoidea) in the Turkish Republic of Northern Cyprus. Turkish Journal of Zoology. 34 (2), 219-224. http://journals.tubitak.gov.tr/zoology/

Instituto Horus, 2011. Alien Invasive Species: Fact Sheets. In: Alien Invasive Species: Fact Sheets. Brazil: http://www.institutohorus.br/inf_fichas_eng.htm

Ion M, Raluca S, 2019. Halyomorpha halys Stål (Heteroptera: Pentatomidae) a new pest for the Oltenia area. Annals of the University of Craiova - Agriculture, Montanology, Cadastre Series. 49 (1), 238-243. http://anale.agro-craiova.ro/index.php/aamc/article/view/946/894

Keçe A F Ç, Ulusoy M R, 2017. Armored scale insects (Hemiptera: Sternorrhyncha: Diaspididae) on ornamental plants in Adana, Turkey. Türkiye Entomoloji Dergisi. 41 (3), 333-346. https://dergipark.org.tr/download/article-file/354688

Kirsur M V, Sakthivel N, Mahimasanthi A, Balasaraswathi S, Bindroo B B, 2014. Weed hosts of papaya mealy bug (Paracoccus marginatus) in mulberry ecosystem of Tamil Nadu. Indian Journal of Sericulture. 53 (1), 78-80.

Lushaj B M, Woodward S, Keča N, Intini M, 2010. Distribution, ecology and host range of Armillaria species in Albania. Forest Pathology. 40 (6), 485-499. DOI:10.1111/j.1439-0329.2009.00624.x

Mallikarjuna B, Magadum S B, Gunashekar V, 2010. A survey on incidence of root diseases of mulberry. Karnataka Journal of Agricultural Sciences. 23 (4), 655.

Mazhar M A, Bagherian S A A, Izadpanah K, 2014. Variants of Hop stunt viroid associated with mulberry vein clearing in Iran. Journal of Phytopathology. 162 (4), 269-271. http://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1439-0434 DOI:10.1111/jph.12182

Meng J R, Liu P P, Zou C W, Wang Z Q, Liao Y M, Cai J H, Qin B X, Chen B S, 2013. First report of a Tospovirus in mulberry. Plant Disease. 97 (7), 1001. DOI:10.1094/PDIS-08-12-0792-PDN

Mohammed E, Ülgentürk S, Uygun N, Garonna A P, Szentkiralayi F, Fent M, Hayat M, 2016. The distribution, host plants and natural enemies of white peach scale, Pseudaulacaspis pentagona (Targioni-Tozzetti) (Hemiptera: Diaspididae), in Ankara Province. Munis Entomology & Zoology. 11 (2), 650-656. http://www.munisentzool.org

Mubashir Niaz, Abbas S Q, Rubab Ayesha, Iftikhar Ali, Tehreema Iftikhar, 2010. New fungal records on Morus alba from Faisalabad Pakistan II. Pakistan Journal of Botany. 42 (3), 1949-1958. http://www.pakbs.org/pjbot/PDFs/42(3)/PJB42(3)1949.pdf

Pane A, Granata G, Cacciola S O, Puglisi I, Evoli M, Aloi F, Spada F la, San Lio G M di, Zambounis A, 2017. First report of root rot of white mulberry caused by simultaneous infections of Phytophthora megasperma and P. multivora in Italy. Plant Disease. 101 (1), 260-261. http://apsjournals.apsnet.org/loi/pdis DOI:10.1094/PDIS-07-16-0993-PDN

Pencheva A, Yovkova M, 2016. New data on alien insect pests of ornamental plants in Bulgaria. Forestry Ideas. 22 (1), 17-33. http://forestry-ideas.info/issues/issues_Index.php?pageNum_rsIssue=1&totalRows_rsIssue=9&journalFilter=55

Peraza-Padilla W, 2014. First report of two Mesocriconema (Nematoda: Criconematidae) species in mulberry trees in Costa Rica. Pesquisa Agropecuária Tropical. 44 (2), 223-229. DOI:10.1590/S1983-40632014000200016

Pieczul K, Jajor E, Perek A, Świerczyńska I, 2017. First report on Mycosphaerella mori on Morus nigra and M. rubra in Poland. Journal of Plant Pathology. 99 (1), 300. http://www.sipav.org/main/jpp/index.php/jpp/article/view/3843/2485

Rashtra Vardhana, 2017. Plant's diseases of district Ghaziabad and adjacent areas. Plant Archives. 17 (1), 727-732. http://www.plantarchives.org/PDF%2017-1/727-732%20(3511).pdf

Rasthra Vardhana, 2007. Plant's havoc by Cuscuta spp. in district Meerut U.P. India. Plant Archives. 7 (2), 917-918.

Rasthra Vardhana, 2007a. Plant's havoc by Cuscuta spp. in district Ghaziabad U.P. India. Plant Archives. 7 (2), 921-922.

Sakthivel P, Karuppuchamy P, Kalyanasundaram M, Srinivasan T, 2012. Host plants of invasive papaya mealybug, Paracoccus marginatus (Williams and Granara de Willink) in Tamil Nadu. Madras Agricultural Journal. 99 (7/9), 615-619. https://doc-00-7g-docsviewer.googleusercontent.com/viewer/securedownload/dsn1aovipa7l846lsfcf94nedj8q2p4u/qo3phtufamvk9q39umu888pbj4t4kkc6/1348647300000/c2l0ZXM=/AGZ5hq8BgbJY1gwaOYx83cPOdNw6/WkdWbVlYVnNkR1J2YldGcGJud3hNWFJvWlcxaFpISmhjMkZuY21samRXeDBkWEpoYkdwdmRYSnVZV3g4WjNnNk56WmpPREk1WXpBd01XWTNZelZrWkE=?a=gp&filename=99-7-9-615-619.pdf&chan=EQAAAOqeu1nfMdjbyOfMSElqQCfRbAOx1kCMBqnRUfeLUnjy&docid=0508176bd4abbdc3e7017b1a89751bc3%7C9c9df36583445f1fe402a841b5e1963b&sec=AHSqidZmGWqJKVKwfKsaqtFstCH

Seebens H, Blackburn T M, Dyer E E, Genovesi P, Hulme P E, Jeschke J M, Pagad S, Pyšek P, Winter M, Arianoutsou M, Bacher S, Blasius B, Brundu G, Capinha C, Celesti-Grapow L, Dawson W, Dullinger S, Fuentes N, Jäger H, Kartesz J, Kenis M, Kreft H, Kühn I, Lenzner B, Liebhold A, Mosena A (et al), 2017. No saturation in the accumulation of alien species worldwide. Nature Communications. 8 (2), 14435. http://www.nature.com/articles/ncomms14435

Shokoohi E, Parastar Z, Panahi H, Abbaspour S, Fourie H, Marais M, 2016. First report of Meloidogyne hispanica in Iran. Australasian Plant Disease Notes. 11 (1), 16. http://link.springer.com/article/10.1007/s13314-016-0202-9

Skelton A, Fowkes A, Adams I, Buxton-Kirk A, Harju V, Forde S, Ward R, Kelly M, Barber P, Fox A, 2018. First report of Prunus necrotic ringspot virus and Mulberry cryptic virus 1 in mulberry (Morus alba) in the United Kingdom. New Disease Reports. 23. DOI:10.5197/j.2044-0588.2018.037.023

Szita É, Fetykó K G, Benedicty Z K, Kozár F, Partsinevelos G, Milonas P, Kaydan M B, 2017. Data on the scale insect (Hemiptera: Coccomorpha) fauna of Greece, with description of two new species. Zootaxa. 4329 (5), 463-476. http://www.mapress.com/j/zt/article/view/zootaxa.4329.5.4 DOI:10.11646/zootaxa.4329.5.4

Tanga C M, Ekesi S, Govender P, Mohamed S A, 2016. Host-plant relationships and natural enemies of the invasive mealybug, Rastrococcus iceryoides Green in Kenya and Tanzania. Journal of Applied Entomology. 140 (9), 655-668. http://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1439-0418 DOI:10.1111/jen.12292

Tunç İ, Bahșİ Ü, Göçmen H, 2012. Thysanoptera fauna of the aegean region, Turkey, in the spring. Turkish Journal of Zoology. 36 (5), 592-606. http://journals.tubitak.gov.tr/zoology/

Wang JiRui, Song ZaoQin, Du YuZhou, 2014. Six new record species of whiteflies (Hemiptera: Aleyrodidae) infesting Morus alba in China. Journal of Insect Science (Annapolis). 14 (136), 1-5. http://www.bioone.org/doi/full/10.1093/jisesa/ieu136

Xie H H, Wei J G, Liu F, Pan X H, Yang X B, 2014. First report of mulberry root rot caused by Lasiodiplodia theobromae in China. Plant Disease. 98 (11), 1581-1582. http://apsjournals.apsnet.org/loi/pdis DOI:10.1094/PDIS-03-14-0261-PDN

Zhang L, Li Y H, Zhang X M, Zhang Q H, Xian H Q, 2018. First report of sour rot caused by Geotrichum candidum on Mori fructus in China. Plant Disease. 102 (12), 2640-2641. DOI:10.1094/PDIS-03-18-0536-PDN

Links to Websites

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GISD/IASPMR: Invasive Alien Species Pathway Management Resource and DAISIE European Invasive Alien Species Gatewayhttps://doi.org/10.5061/dryad.m93f6Data source for updated system data added to species habitat list.
Global register of Introduced and Invasive species (GRIIS)http://griis.org/Data source for updated system data added to species habitat list.

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