Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide


Marasmia exigua
(rice leaf roller)



Marasmia exigua (rice leaf roller)


  • Last modified
  • 14 July 2018
  • Datasheet Type(s)
  • Pest
  • Natural Enemy
  • Preferred Scientific Name
  • Marasmia exigua
  • Preferred Common Name
  • rice leaf roller
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Metazoa
  •     Phylum: Arthropoda
  •       Subphylum: Uniramia
  •         Class: Insecta

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TitleM. exigua; adult (museum set specimen)
Copyright©David Agassiz
M. exigua; adult (museum set specimen)©David Agassiz


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Preferred Scientific Name

  • Marasmia exigua (Butler 1879)

Preferred Common Name

  • rice leaf roller

Other Scientific Names

  • Cnaphalocrocis exigua (Butler, 1879)
  • Samea exigua Butler 1879
  • Susumia exigua (Butler); Marumo 1930
  • Susumia nawae Matsumura, 1920

International Common Names

  • English: rice, leafroller
  • French: tordeuse du riz

Local Common Names

  • Japan: Ine-tatehamaki

EPPO code

  • SUSUEX (Susumia exigua)

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta
  •                     Order: Lepidoptera
  •                         Family: Crambidae
  •                             Genus: Marasmia
  •                                 Species: Marasmia exigua

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Growth Stages

Top of page Flowering stage, Seedling stage, Vegetative growing stage

List of Symptoms/Signs

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SignLife StagesType
Leaves / external feeding
Leaves / leaves rolled or folded

Natural enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Apanteles guamensis Parasite Larvae
Aulacocentrum philippinense Parasite Larvae
Bacillus thuringiensis thuringiensis Pathogen Larvae
Beauveria bassiana Pathogen Larvae
Cardiochiles philippinensis Parasite Larvae
Coccigomimus nipponicus Parasite Pupae
Copidosomopsis nacoleiae Parasite Eggs/Larvae
Cotesia Parasite Larvae
Erigonidium graminicolum Predator China; Hunan rice
Erynia radicans Pathogen Larvae
Goniozus nr. triangulifer Parasite Larvae
Ischnojoppa luteator Parasite Larvae
Itoplectis naranyae Parasite Larvae
Lycosa pseudoannulata Predator
Nomuraea rileyi Pathogen Larvae
Nucleopolyhedrosis virus Larvae
Orgilus Parasite Larvae
Paecilomyces farinosus Pathogen Larvae
Phaeogenes Parasite Larvae
Pimpla nipponicus Parasite
Temelucha philippinensis Parasite Larvae India; West Bengal rice
Temelucha stangli Parasite Larvae
Trathala flavoorbitalis Parasite Larvae Fiji rice
Trichogramma Parasite Eggs
Trichogramma chilotraeae Parasite Eggs
Trichogramma evanescens Parasite Eggs
Trichogrammatoidea australicum Parasite Eggs
Trichomma cnaphalocrocis Parasite Larvae
Tropobracon schoenobii Parasite Larvae
Vulgichneumon diminutus Parasite Pupae


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APPPC, 1987. Insect pests of economic significance affecting major crops of the countries in Asia and the Pacific region. Technical Document No. 135. Bangkok, Thailand: Regional Office for Asia and the Pacific region (RAPA).

Barrion AT; Litsinger JA, 1985. Identification of rice leaffolders (LF) by wing markings. International Rice Research Newsletter, 10(1):24

CIE, 1977. Distribution maps of pests: Susumia exigua, Map 365. Wallingford, UK: CAB International

Dale D, 1994. Insect pests of the rice plant and their biology and ecology. In: Heinrichs EA, ed. Biology and Management of Rice Insects. Wiley Eastern Limited and International Rice Research Institute.

Grist DH; Lever RJAW, 1969. Pests of Rice. London, UK: Longman.

Hinckley AD, 1963. The rice leafroller, Susumia exigua (Butler) in Fiji. Journal of Economic Entomology, 56:112-113.

Hupeh Academy of Agricultural Science (HAAS), 1978. The Insect Pests of Rice and Their Natural Enemies. China: Wuchang.

Khan ZR; Barrion AT; Litsinger JA; Castilla NP; Joshi RC, 1988. A bibliography of rice leaffolders (Lepidoptera: Pyralidae). Insect Science and its Application, 9(2):129-174

Lee SC; Song YH; Lee JO; Han SC, 1973. Minor pests. In: Literature Review of Korean Rice Pests. Suweon, Korea: Department of Entomology, Institute of Agricultural Science, Office of Rural Development, 30-33.

Lever RJAW, 1946. The rice leafroller, Marasmia venilialis (sic). Agricultural Journal of Fiji, 17:11.

Munroe E, 1991. Transfer of Aulacodes eupselias Meyrick to Pyraustinae, with notes on the genus Marasmia Lederer and on cataclystiform wing patterns in the family Crambidae (Lepidoptera:Pyralidae). Bishop Museum Occasional Papers, 31:122-130.

Distribution Maps

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