Invasive Species Compendium

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Datasheet

Luffa aegyptiaca
(loofah)

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Datasheet

Luffa aegyptiaca (loofah)

Summary

  • Last modified
  • 18 November 2019
  • Datasheet Type(s)
  • Invasive Species
  • Host Plant
  • Preferred Scientific Name
  • Luffa aegyptiaca
  • Preferred Common Name
  • loofah
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Plantae
  •     Phylum: Spermatophyta
  •       Subphylum: Angiospermae
  •         Class: Dicotyledonae
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    UK
    compend@cabi.org
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Identity

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Preferred Scientific Name

  • Luffa aegyptiaca P. Miller

Preferred Common Name

  • loofah

Other Scientific Names

  • Cucumis lineatus Bosc
  • Cucurbita luffa hort.
  • Luffa cathu-picinna Ser.
  • Luffa clavata Roxb.
  • Luffa cylindrica (L.) M. J. Roem.
  • Luffa insularum A. Gray
  • Luffa leiocarpa F. Muell.
  • Luffa leucosperma M. J. Roem.
  • Luffa pentandra Roxb.
  • Luffa petola Ser.
  • Luffa racemosa Roxb.
  • Luffa scabra Schumach.
  • Luffa striata Schrad.
  • Luffa veitchii Naud.
  • Momordica cylindrica L.
  • Momordica luffa L.
  • Momordica operculata Blanco
  • Turia cordata J. F. Gmel.

International Common Names

  • English: climbing okra; dish-cloth gourd; dishrag gourd; luffa; rag gourd; smooth loofah; sponge gourd; sponge luffa; vegetable sponge
  • Spanish: esponja vegetal; estropajo; lufa; pashte (Mexico); paste; pepino para paste
  • French: courge cylindrique de chine; courge torchon; eponge végétale; luffa d'Egypte; pétole; torchon
  • Arabic: lûf
  • Chinese: shui gua; shui kwa; si gua; tian luo gua; tian si gua
  • Portuguese: bucha

Local Common Names

  • Germany: Ågyptische Schwammkürbis; Schwammgurke; Schwammkuerbis; Schwammkürbis
  • India: dhundal; jhinga; meethi torai; mozhuka peerkankai; turai
  • Indonesia: belustru; blustru
  • Malaysia: ketola manis; petola buntal; petola manis
  • Nepal: ghiu toriya; palo
  • Netherlands: komkommer, spons-
  • Sri Lanka: vatakolu
  • Sudan: emes
  • Thailand: boap hom; buap hom; buap klom

EPPO code

  • LUFAE (Luffa aegyptiaca)

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Plantae
  •         Phylum: Spermatophyta
  •             Subphylum: Angiospermae
  •                 Class: Dicotyledonae
  •                     Order: Violales
  •                         Family: Cucurbitaceae
  •                             Genus: Luffa
  •                                 Species: Luffa aegyptiaca

Notes on Taxonomy and Nomenclature

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The botanical name Luffa aegyptica is due to Phillip Miller who first saw the gourd growing in Egyptian gardens in the 17th century.

Description

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Sponge gourd is a vigorous-growing vine comprising five angled stems with tendrils on the axils of leaves to facilitate climbing. The leaves are broadly ovate to reniform, dark green with silvery patches on the topside, 5–7 deep lobes, dentate, scabrous, 6–25 x 8–27 cm, apex acute, base cordate, petiole hispid and 5–10 cm long (Purseglove, 1976). The flowers are monoecious, produced on leaf axils, and consist of five petals of a conspicuous yellow colour, 5–10 cm across, united below in a bell-shaped corolla, open in early morning. The staminate flowers are produced in racemes of 4–20 and contain five free stamens. The pistillate flowers are borne either singly on a separate node or in association with staminate inflorescence. The lower nodes usually bear only staminate inflorescence, middle nodes both staminate inflorescence and pistillate flowers, and terminal nodes only pistillate flowers (Omini and Hossain, 1987). The female flowers have three stigmas and a smooth, cylindrical ovary. The fruits are green with longitudinal marked lines, up to 61 cm long and 7.6 cm in diameter, shaped like a large cylindrical pepo (berry). The endocarp is the edible portion of the fruit that later becomes fibrous and spongy. The seeds are borne with parietal placentation. The dry, mature fruit is composed of an outer hard shell and an inner dense network of cellulose fibre (sponge) containing hundreds of flat, smooth, black seeds (Shah et al., 1980). The fruit is essentially an inferior berry, even though it is called a pepo because of its hard, tough rind when mature (Seshadri, 1999).

Distribution

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L. aegyptica is agreed to be of tropical origin but the geographical region of origin is not certain. The crop has been under cultivation since ancient times. It is widely believed to have been domesticated in tropical Asia, possibly India, and reached China about 600 AD (Purseglove, 1976) and Egypt in the Middle Ages (Mansfeld, 2001). It has been cultivated in the Middle East, India, China, Japan and Malaysia for centuries (Porterfield, 1955). The word luffa or 'loofah' is of Arabic origin and the sponge characteristic of the crop has been described in old Egyptian writings (Seshadri, 1999). Sponge gourd is now found in cultivation or as an escape in all tropical regions of the world. It was an important crop used for industrial filters until World War II and Japan cultivated the best crop for this purpose.The main commercial production areas today are China, Korea, India, Japan, Central America and Brazil.

Cultivation of sponge gourd has been reported in the following countries: Algeria, Argentina, Australia, Belize, Benin, Bhutan, Bolivia, Brazil, Burkina Faso, Burundi, Cameroon, Chad, China, Colombia, Comoros, Costa Rica, Côte d'Ivoire, Ecuador, El Salvador, Egypt, Equatorial Guinea, Ethiopia, French Polynesia, Georgia, Germany, Ghana, Guatemala, Guinea, Guyana, Honduras, India, Iraq, Italy, Jamaica, Japan, Kenya, Kyrgyzstan, Laos, Liberia, Libya, Madagascar, Mali, Myanmar, Mexico, Nepal, Nicaragua, Nigeria, Pakistan, Panama, Papua New Guinea, Paraguay, Peru, Philippines, Portugal, Russian Federation, Senegal, South Africa, Spain, Sudan, Suriname, Switzerland, Tajikistan, Tanzania, Thailand, Trinidad, Tunisia, Uganda, Ukraine, UK, USA, Uzbekistan, Venezuela and Vietnam.

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 25 Feb 2021
Continent/Country/Region Distribution Last Reported Origin First Reported Invasive Reference Notes

Africa

AlgeriaPresent
BeninPresent
Burkina FasoPresent
BurundiPresent
CameroonPresent
ChadPresent
ComorosPresent
Côte d'IvoirePresent
EgyptPresent
Equatorial GuineaPresent
EthiopiaPresent
GambiaPresent
GhanaPresent
GuineaPresent
KenyaPresent
LiberiaPresent
LibyaPresent
MadagascarPresent
MaliPresent
MayottePresent
NigerPresent
NigeriaPresent
SenegalPresent
South AfricaPresent
SudanPresent
TanzaniaPresent
TunisiaPresent
UgandaPresent

Asia

BhutanPresent
ChinaPresent
-LiaoningPresent
-SichuanPresent
GeorgiaPresent
IndiaPresent
-ChhattisgarhPresent
-PunjabPresent
-Uttar PradeshPresent
IndonesiaPresent
IraqPresent
JapanPresent
KyrgyzstanPresent
LaosPresent
MyanmarPresent
NepalPresent
PakistanPresent
PhilippinesPresent
South KoreaPresent
Sri LankaPresent
TaiwanPresentOriginal citation: Fang Chang (1984)
TajikistanPresent
ThailandPresent
UzbekistanPresent
VietnamPresent
YemenPresent

Europe

GermanyPresent
ItalyPresent
PortugalPresent
RussiaPresent
SpainPresent
SwitzerlandPresent
UkrainePresent
United KingdomPresent

North America

BelizePresent
Costa RicaPresent
El SalvadorPresent
GrenadaPresent
GuatemalaPresent
HondurasPresent
JamaicaPresent
MexicoPresent
NicaraguaPresent
PanamaPresent
Trinidad and TobagoPresent
United StatesPresent
-FloridaPresentOriginal citation: USDA, NRCS (2007)
-IllinoisPresentOriginal citation: USDA, NRCS (2007)
-LouisianaPresentOriginal citation: USDA, NRCS (2007)
-North CarolinaPresent
-VirginiaPresentOriginal citation: USDA, NRCS (2007)

Oceania

American SamoaPresent
AustraliaPresent
Christmas IslandPresent
Cook IslandsPresent
Federated States of MicronesiaPresentChuuk, Pohnpei and Kosrae
FijiPresent
French PolynesiaPresent, Few occurrencesTahiti
GuamPresent
Marshall IslandsPresent
NauruPresent
New CaledoniaPresent
NiuePresent
Northern Mariana IslandsPresent
PalauPresent
Papua New GuineaPresent
SamoaPresent
Solomon IslandsPresent
TongaPresent
Wallis and FutunaPresent

South America

ArgentinaPresent
BoliviaPresent
BrazilPresent
-BahiaPresent
-Minas GeraisPresent
-Rio Grande do SulPresent
ColombiaPresent
EcuadorPresent
GuyanaPresent
ParaguayPresent
PeruPresent
SurinamePresent
VenezuelaPresent

Biology and Ecology

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Sponge gourd has an annual growth habit. It is propagated by seeds obtained from ripe fruits. The plant is a vine with tendrils bearing monoecious flowers. The pistillate flowers are pollinated by insects. Although sponge gourd is considered a cross pollinated crop, the absence of self-incompatibility permits natural self pollination taking place within the same plant. As insects do not differentiate flowers from the same or different plants, 20–40% self pollination can be expected. To ensure the visit of pollinating insects, the flowers are showy in colour, large in size and staminate flowers are produced in greater abundance than pistillate ones (Seshadri, 1999). Smooth luffa produces fruits that are grouped as pepo. Each fruit contains hundreds of flat, smooth, black seeds.

Sponge gourd tolerates a wide range of climatic and soil conditions. It is adapted to tropical and subtropical climates and requires warm summer temperatures and a long, frost-free growing season for maximum yield. The average temperature should be around 30–35°C. It grows best at altitudes below 500 m. Sponge gourd seeds require warm soil to germinate. The growing season can be extended by using transplants grown under glass earlier in the season. Well distributed rainfall during the growing season is needed for optimum crop growth and both growth and yield are substantially reduced under water stress. Soil moisture of at least 10–15% above the permanent wilting point is required to maintain growth (Seshadri, 1999). Excessive rainfall during the flowering and fruiting period can damage yield and lower fruit quality (Tindall, 1983). Sponge gourd will grow on many soil types but well-drained sandy loams are preferred. It is sensitive to acid soils; the optimal soil pH is 6.0–6.8.

Means of Movement and Dispersal

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L. aegyptiaca is spread by seed, which is dispersed by birds and rodents and also planted by humans.

Impact

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From Englberger (2009):

'This fast-growing vine, with its smothering growth, can be very aggressive. The plant is often found on fences and overgrows native vegetation. It is usually found in open, disturbed areas like roadsides.'

Uses

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Sponge gourd fruit is consumed as a vegetable when young in the Indian subcontinent, South-East Asia and the Caribbean. It can be eaten as a curried vegetable, used in soup, fried, boiled, stuffed, dried and pickled. The fruit is cut into small pieces, like squash, for cooking. Fried flower buds and flowers are delicacies in Indian and Chinese cuisine and raw flowers are a nice addition to tossed salads. The fruit contains 93% water, 1.2% protein, 0.2% fat, 3.1% carbohydrate, 2.0% fibre and 0.5% ash (Purseglove, 1976). Du et al. (2006) reported that luffa fruit was a good source of antioxidants. The seed yields edible oil. Seed kernels, approximately 51% weight of the seeds, contain 46% oil and 40% protein (Purseglove, 1976).

Different plant parts are used in native medicines in India, China and in other countries. Cucurbits are a well recognized source of secondary metabolites. Ng et al. (1992) reported two proteins from seeds of sponge gourd that had the capability to induce mid-term abortion in mice, inhibit protein synthesis in a cell-free system, and suppress thymidine uptake in human choriocarcinoma cells. Ng (1993) found a mention of the use of cucurbits to induce second trimester abortions in China dating back to 1920. Sponge gourd is also used in Chinese medicine to treat rheumatic pain. El-Fiky et al. (1996) observed a significant decrease in the blood sugar of streptozotocin diabetic rats by the oral administration of ethanolic extracts of L. aegyptiaca seeds, while producing no changes in blood sugar levels in normal rats. Sap from the stem is used in toilet preparations in Japan.

Portuguese explorers first discovered uses for the fibro-vascular network of the ripe fruit of sponge gourd. It is presently used as bath sponge, pot scrubber, filter, packing material and for making handicrafts (Porterfield, 1955; Heiser, 1979). Fibre also possesses sound- and shock-absorbing properties and is therefore used in sandals, insoles, gloves and helmets. Sponges of different fibre density are needed for different products (Davis and DeCourley, 1993). The largest market for luffa sponge is in the cosmetic industry, where it is used in various bath and cosmetic products.

Luffa sponge has been found to be effective in the removal of heavy metal from aqueous solutions. Iqbal and Edyvean (2004) found that alginate-coated loofa sponge removed cadmium, rapidly reaching an equilibrium loading of 124 mg/g in 30 min with 70% of equilibrium uptake achieved in 10 min. In contrast, it took 240 min for alginate beads to reach a loading equilibrium of 88 mg/g under identical conditions.

Uses List

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Human food and beverage

  • Fruits
  • Oil/fat
  • Vegetable

Materials

  • Fibre

Medicinal, pharmaceutical

  • Traditional/folklore

Prevention and Control

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Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.

From Englberger (2009):

'Physical control can be effective if used repeatedly for and extended period. For chemical control, triclopyr and glyphosate can be used.'

References

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Davis JM, 1994. Luffa sponge gourd production practices for temperate climates. HortScience, 29:263-266

Davis JM, DeCourley CD, 1993. Luffa sponge gourds: a potential crop for small farms. In: Janick J, Simon JE, eds. New Crops. New York, USA: Wiley, 560-561

DOASL, 2006. Luffa. Department of Agriculture, Sri Lanka. Online at: http://www.agridept.gov.lk/agri_tech_search.php?hid=Vegetables&shid=Luffa&mhid=Crop%20 management, modified February 2006

Du Q, Jerg G, Winterhalter P, Zhao Y, Xu Y, Li L, 2006. Antioxidant constituents in the fruits of Luffa cylindrica (L.) Roem. Journal of Agricultural and Food Chemistry, 54:4186-4190

El-Fiky FK, Abou-Karam MA, Afify EA, 1996. Effect of Luffa aegyptiaca (seeds) and Carissa edulis (leaves) extracts on blood glucose level of normal and streptozotocin diabetic rats. Journal Ethnopharmacology, 50:43-7

Ellington TL, Wehner TC, 1996. Post-harvest treatments for producing sponges from immature fruits of luffa gourd. Cucurbit Genetic Coop. Report, 19:85-86

Englberger K, 2009. Invasive weeds of Pohnpei: A guide for identification and public awareness. Kolonia, Federated States of Micronesia: Conservation Society of Pohnpei, 29 pp

Esquinas-Alcazar JT, Gulick PJ, 1983. Genetic resources of Cucurbitaceae. Rome, Italy: International Board for Plant Genetic Resource

Fang MN, Chang CP, 1984. The injury and seasonal occurrence of melon fly, Dacus cucurbitae Coquillett, in central Taiwan (Trypetidae, Diptera). Plant Protection Bulletin, Taiwan, 26(3):241-248

Heiser CB, 1979. The Gourd Book. Norman, USA: University of Oklahoma Press

Iqbal M, Edyvean RGJ, 2004. Alginate coated loofa sponge discs for the removal of cadmium from aqueous solutions. Biotechnology Letters, 26:165-169

Malik IJ, Ellington TL, Wehner TC, Sanders DC, 2001. Seed treatment effects on emergence of Luffa sponge gourd. Cucurbit Genetic Coop. Report, 24:107-109

Mansfeld R, 2001. Mansfeld’s encyclopedia of agricultural and horticultural crops. Berlin, Germany: Springer

Ng TB, Wong RNS, Yeung HW, 1992. Two proteins with ribosome-inactivating, cytotoxic and abortifacient activities from seeds of Luffa cylindrica Roem (Cucurbitaceae). Biochemistry International, 27:197-207

Ng TJ, 1993. New opportunities in Cucurbitaceae. In: Janick J, Simon JE, eds. New Crops. New York, USA: Wiley, 538-546

Okusanya OT, Ola Adams BA, Bamidele JF, 1981. Variation in size, leaf morphology and fruit characters among 25 populations of Luffa aegyptica. Canadian Journal Botany, 59:2618-2627

Omini ME, Hossain MG, 1987. Modification of sex expression in sponge gourd (Luffa cylindrica L. Roem) by mineral nutrient treatments. Genetica, 74:203-209

Pathak GN, Singh SN, 1949. Studies in genus Luffa. I. Cytogenetic investigations in the interspecific hybrid L. cylindrica x L. acutangula. Indian Journal Genetics Plant Breeding, 9:18-26

PIER, 2007. Pacific Island Ecosystems at Risk (PIER). Institute of Pacific Islands Forestry. http://www.hear.org/pier/species/luffa_aegyptiaca.htm

Porterfield WM, 1955. Loofah: the sponge gourd. Economic Botany, 9:211-223

Purseglove JW, 1976. Tropical Crops: Dicotyledons. London, UK: Longman Group

Sahni GP, Singh RK, Saha BC, 1987. Genotypic and phenotypic variability in ridge gourd. Indian Journal of Agricultural Sciences, 57:666-668

Seshadri VS, 1999. Cucurbits. In: Bose TK, Som MG, Kabir J, eds. Vegetable Crops. Calcutta, India: Naya Prokash, 91-164

Shah JJ, Thanki YJ, Kothari IL, 1980. Skeletal fibrous net in fruits of Luffa cylindrica M. Roem. and Luffa acutangula Roxb. In: Nagaraj M, Malik CP, eds. Current Trends in Botanical Research. New Delhi, India: Kalyani Publishers, 61-72

Singh PK, Dasgupta SK, 2004. Hybrid luffah. Journal of New Seeds, 6:211-215

Sinnott EW, Bloch R, 1943. Luffa sponges, a new crop for the Americas. Journal New York Botanical Garden, 44:125-132

Talpur MA, Hussain T, Rustamani MA, Siddiqui MD, Chhutto AB, 1998. Comparative bio-efficacy of baits and insecticides in controlling Ethiopian melon fly, Bactrocera ciliatus (Loew) on sponge gourd. Pakistan Entomologist, 20(1/2): 84-86

Tindall HD, 1983. Vegetables in the tropics. London, United Kingdom; Macmillan Press; Westport, Connecticut, USA; AVI Technical Books, xi + 533 pp

USDA, NRCS, 2007. The PLANTS Database. Baton Rouge, USA: National Plant Data Center. http://plants.usda.gov/

Wehner TC, Ellington TL, 1997. Seed treatment effects on emergence of luffa sponge gourd. Cucurbit Genetic Coop. Report, 20:63-64

Wehner TC, Ellington TL, 1999. Vegetable cultivar descriptions for North America, list 24, Gourd-Luffa Sponge. HortScience, 34:800

Zipcode Zoo, 2007. Luffa aegyptiaca. http://zipcodezoo.com/Plants/L/Luffa_aegyptiaca/. BayScience Foundation, Inc

Distribution References

Abdel-Salam A M M, Rezk A A, Dawoud R A, 2019. Biochemical, serological, molecular and natural host studies on Tomato Chlorosis Virus in Egypt. Pakistan Journal of Biological Sciences. 22 (2), 83-94. https://scialert.net/fulltext/?doi=pjbs.2019.83.94&org=11

Anwar S A, McKenry M V, 2010. Incidence and reproduction of Meloidogyne incognita on vegetable crop genotypes. Pakistan Journal of Zoology. 42 (2), 135-141.

Badii K B, Billah M K, Afreh-Nuamah K, Obeng-Ofori D, 2015. Species composition and host range of fruit-infesting flies (Diptera: Tephritidae) in northern Ghana. International Journal of Tropical Insect Science. 35 (3), 137-151. http://journals.cambridge.org/action/displayJournal?jid=JTI

Bellé C, Moccellin R, Meneses P R, Neves C G, Groth M Z, Kaspary T E, Barros D R de, Farias C R J de, 2018. First report of Sclerotium rolfsii causing stem rot of Luffa cylindrica in Brazil. Plant Disease. 102 (1), 250-251. DOI:10.1094/PDIS-07-17-1073-PDN

CABI, Undated. Compendium record. Wallingford, UK: CABI

CABI, Undated a. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI

Cho S E, Hong S H, Kim B S, Kim J Y, Shin H D, 2015. First report of powdery mildew caused by Podosphaera xanthii on Luffa cylindrica in Korea. Plant Disease. 99 (12), 1861. http://apsjournals.apsnet.org/loi/pdis DOI:10.1094/PDIS-05-15-0518-PDN

Englberger K, 2009. Invasive weeds of Pohnpei: a guide for identification and public awareness. In: Invasive weeds of Pohnpei: a guide for identification and public awareness, Kolonia, Federated States of Micronesia: Conservation Society of Pohnpei (CSP). 29 pp.

Falak Naz, Mian Inayatullah, 2013. Henosepilachna septima dieke (Coccinellidae; Coleoptera); a new record for Pakistan along with notes on its taxonomy, host plants and distribution. Sarhad Journal of Agriculture. 29 (2), 235-238. http://www.aup.edu.pk/sj_pdf/10%20407-2012%20HENOSEPILACHNA%20SEPTIMA%20DIEKE%20(COCCINELLIDAE_%20COLEOPTERA)_%20A%20NEW%20RECORD%20FOR%20PAKISTAN%20ALONG%20WITH%20NOTES%20ON%20ITS%20TAXONOMY,%20HOST%20PLANTS%20AND%20DISTRIBUTION.pdf

Li B J, Li H L, Shi Y X, Xie X W, 2014. First report of Pseudomonas cichorii causing leaf spot of vegetable sponge gourd in China. Plant Disease. 98 (1), 153. http://apsjournals.apsnet.org/loi/pdis DOI:10.1094/PDIS-03-13-0312-PDN

Liu Ning, Zhou XiaoSong, Chen LiJie, Duan YuXi, 2013. Description of a new Pratylenchus species from China (Tylenchida, Pratylenchidae). Zoological Research. 34 (5), E140-E142. http://www.zoores.ac.cn/EN/abstract/abstract3398.shtml

Mware B, Olubayo F, Narla R, Songa J, Amata R, Kyamanywa S, Ateka E M, 2010. First record of spiraling whitefly in coastal Kenya: emergence, host range, distribution and association with cassava brown streak virus disease. International Journal of Agriculture and Biology. 12 (3), 411-415. http://www.fspublishers.org/ijab/past-issues/IJABVOL_12_NO_3/18.pdf

Patil K P, Awadhiya G K, Pandey S R, 2017. Occurrence of powdery mildew on some plants from Raipur of Chhattisgarh state. Trends in Biosciences. 10 (32), 6818-6829. http://trendsinbiosciencesjournal.com/upload/23-8868_(K_P__Patila).pdf

PIER, 2007. Pacific Island Ecosystems at Risk (PIER). In: Institute of Pacific Islands Forestry, http://www.hear.org/pier/species/luffa_aegyptiaca.htm

Raj Verma, Baranwal V K, Satya Prakash, Tomer S P S, Pant R P, Ahlawat Y S, 2006. First report of Papaya ringspot virus W in sponge gourd from India. Plant Disease. 90 (7), 974. DOI:10.1094/PD-90-0974B

Rashtra Vardhana, 2017. Plant's diseases of district Ghaziabad and adjacent areas. Plant Archives. 17 (1), 727-732. http://www.plantarchives.org/PDF%2017-1/727-732%20(3511).pdf

Shimizu Y, Kohama T, Uesato T, Matsuyama T, Yamagishi M, 2007. Invasion of solanum fruit fly Bactrocera latifrons (Diptera: Tephritidae) to Yonaguni Island, Okinawa Prefecture, Japan. Applied Entomology and Zoology. 42 (2), 269-275. http://odokon.ac.affrc.go.jp/ DOI:10.1303/aez.2007.269

Silva M, Freitas N M, Mendonça H L, Barreto R W, 2013. First report of Stagonosporiopsis cucurbitacearum causing fruit rot of Luffa cylindrica in Brazil. Plant Disease. 97 (8), 1120-1121. DOI:10.1094/PDIS-02-13-0175-PDN

Talpur M A, Hussain T, Rustamani M A, Siddiqui M D, Chhutto A B, 1998. Comparative bio-efficacy of baits and insecticides in controlling Ethiopian melon fly, Bactrocera ciliatus (Loew) on sponge gourd. Pakistan Entomologist. 20 (1/2), 84-86.

Wallace E, Choi Y J, Thines M, Quesada-Ocampo L M, 2016. First report of Plasmopara aff. australis on Luffa cylindrica in the United States. Plant Disease. 100 (2), 537. DOI:10.1094/PDIS-06-15-0684-PDN

Zia-ur-Rehman M, Herrmann H W, Hameed U, Haider M S, Brown J K, 2013. First detection of Cotton leaf curl burewala virus and cognate Cotton leaf curl Multan betasatellite and Gossypium darwinii symptomless alphasatellite in symptomatic Luffa cylindrica in Pakistan. Plant Disease. 97 (8), 1122. http://apsjournals.apsnet.org/loi/pdis DOI:10.1094/PDIS-12-12-1159-PDN

Zipcode Zoo, 2007. Luffa aegyptiaca., BayScience Foundation, Inc. http://zipcodezoo.com/Plants/L/Luffa_aegyptiaca/

Links to Websites

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WebsiteURLComment
GISD/IASPMR: Invasive Alien Species Pathway Management Resource and DAISIE European Invasive Alien Species Gatewayhttps://doi.org/10.5061/dryad.m93f6Data source for updated system data added to species habitat list.

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