Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide


Cydia caryana
(hickory shuckworm)



Cydia caryana (hickory shuckworm)


  • Last modified
  • 10 December 2020
  • Datasheet Type(s)
  • Pest
  • Preferred Scientific Name
  • Cydia caryana
  • Preferred Common Name
  • hickory shuckworm
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Metazoa
  •     Phylum: Arthropoda
  •       Subphylum: Uniramia
  •         Class: Insecta
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Preferred Scientific Name

  • Cydia caryana Fitch

Preferred Common Name

  • hickory shuckworm

Other Scientific Names

  • Enarmonia caryana Fitch
  • Ephippiphora caryana Fitch
  • Grapholitha caryae
  • Grapholitha caryana Fitch
  • Laspeyresia caryana Fitch

International Common Names

  • English: hickory, shuckworm
  • Spanish: gusano de la cascara de la nuez (mexico)
  • French: foreur des brous

EPPO code

  • LASPCA (Cydia caryana)

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta
  •                     Order: Lepidoptera
  •                         Family: Tortricidae
  •                             Genus: Cydia
  •                                 Species: Cydia caryana

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 05 Aug 2021

Natural enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Apanteles epinotiae Parasite Arthropods|Larvae
Bracon cushmani Parasite Arthropods|Larvae
Calliephialtes grapholithae Parasite
Eurytoma pini Parasite Arthropods|Larvae; Arthropods|Pupae
Holepyris floridanus Parasite
Lixophaga mediocris Parasite
Macrocentrus instabilis Parasite
Phanerotoma fasciata Parasite Arthropods|Larvae

Distribution Maps

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