Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide

Datasheet

Eutetranychus orientalis
(Citrus brown mite)

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Datasheet

Eutetranychus orientalis (Citrus brown mite)

Summary

  • Last modified
  • 10 December 2020
  • Datasheet Type(s)
  • Pest
  • Natural Enemy
  • Preferred Scientific Name
  • Eutetranychus orientalis
  • Preferred Common Name
  • Citrus brown mite
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Metazoa
  •     Phylum: Arthropoda
  •       Subphylum: Chelicerata
  •         Class: Arachnida
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    Compendia
    CAB International
    Wallingford
    Oxfordshire
    OX10 8DE
    UK
    compend@cabi.org
  • Distribution map More information

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Identity

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Preferred Scientific Name

  • Eutetranychus orientalis Klein

Preferred Common Name

  • Citrus brown mite

Other Scientific Names

  • Anychus latus
  • Anychus orientalis
  • Anychus ricini
  • Eutetranychus anneckei Meyer
  • Eutetranychus latus
  • Eutetranychus monodi Andre
  • Eutetranychus sudanicus Elbadry

International Common Names

  • English: oriental mite; oriental red mite; oriental spider mite
  • Spanish: acaro rojo oriental
  • French: tétranique rouge asiatique

Local Common Names

  • Germany: Orientalische spinnmilbe
  • Israel: haakarit haaduma hamizrachit
  • South Africa: lowveld citrus mite

EPPO code

  • EUTEAN (Eutetranychus anneckei)
  • EUTELA (Eutetranychus latus)
  • EUTEMO (Eutetranychus monodi)
  • EUTEOR (Eutetranychus orientalis)
  • EUTESU (Eutetranychus sudanicus)

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Chelicerata
  •                 Class: Arachnida
  •                     Subclass: Acari
  •                         Superorder: Acariformes
  •                             Suborder: Prostigmata
  •                                 Family: Tetranychidae
  •                                     Genus: Eutetranychus
  •                                         Species: Eutetranychus orientalis

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 19 May 2021
Continent/Country/Region Distribution Last Reported Origin First Reported Invasive Reference Notes

Africa

Cabo VerdePresent
EgyptPresent
EswatiniPresent
EthiopiaPresent
GambiaAbsent, Unconfirmed presence record(s)
KenyaPresent
LibyaPresent
MalawiPresent
MaliPresent
MauritaniaPresent
MoroccoPresent
MozambiquePresent
NigeriaPresent
SenegalPresent
South AfricaPresent
SudanPresent
TunisiaPresent

Asia

AfghanistanPresent
BangladeshPresent
ChinaPresent
-GuangdongPresent
-GuangxiPresent
-HainanPresent
-ShandongPresent
-SichuanPresent
Hong KongPresent
IndiaPresent
-AssamPresent
-DelhiPresent
-HaryanaPresent
-Himachal PradeshPresent
-Jammu and KashmirPresent
-JharkhandPresent
-KarnatakaPresent
-KeralaPresent
-Madhya PradeshPresent
-MaharashtraPresent
-MeghalayaPresent
-PunjabPresent
-RajasthanPresent
-Tamil NaduPresent
-Uttar PradeshPresent
-West BengalPresent
IndonesiaPresent
-SumatraPresent
IranPresent
IraqPresent
IsraelPresent, Widespread
JapanPresent, Localized
-HonshuPresent
-Ryukyu IslandsPresent
JordanPresent
KuwaitPresent
LebanonPresent
MalaysiaPresent
-Peninsular MalaysiaPresent
NepalPresent
PakistanPresent
PhilippinesAbsent, Invalid presence record(s)
Saudi ArabiaPresent
SyriaPresent
TaiwanPresent
ThailandPresent
TurkeyPresent
United Arab EmiratesPresent
YemenPresent

Europe

AustriaAbsent
BelgiumAbsent
CyprusPresent
GreecePresent
NetherlandsAbsent, Confirmed absent by survey
SerbiaPresentvia PestLens newsletter
SloveniaAbsent
SpainPresent, Localized

Oceania

AustraliaPresent, Localized
-New South WalesPresent
-Northern TerritoryPresent
-QueenslandPresent
-Western AustraliaPresent

Growth Stages

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Vegetative growing stage

List of Symptoms/Signs

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SignLife StagesType
Leaves / abnormal colours
Leaves / external feeding
Leaves / necrotic areas

Natural enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Acaropsellina sollers Predator Adults/Nymphs
Acaropsis aegyptiaca Predator Adults/Nymphs
Agistemus exsertus Predator Adults/Nymphs
Amblyseius alstoniae Predator Adults/Nymphs
Amblyseius herbicolus Predator Adults/Nymphs
Amblyseius maai Predator Adults/Nymphs
Amblyseius okinawanus Predator Adults/Nymphs
Amblyseius rubini Predator Adults/Eggs/Larvae/Nymphs
Amblyseius swirskii Predator Adults/Eggs/Larvae/Nymphs
Amblyseius taiwanicus Predator Adults/Nymphs
Chiapacheylus macrocorneus Parasite
Cunaxa capreolus Predator Adults/Nymphs
Euseius gossipi Predator Adults/Nymphs
Euseius ovalis Predator Adults/Nymphs
Euseius scutalis Predator Adults/Eggs/Larvae/Nymphs
Hirsutella thompsonii Pathogen Adults/Larvae/Nymphs
Lecanicillium lecanii Pathogen Adults/Eggs/Larvae/Nymphs
Metaseiulus occidentalis Predator Adults/Nymphs
Neoseiulus longispinosus Predator Adults/Nymphs
Oligota oviformis Predator Adults/Nymphs
Pronematus ubiquitus Predator Adults/Nymphs
Scolothrips indicus Predator Adults/Nymphs
Scolothrips longicornis Predator Adults/Nymphs
Seiulus finlandicus Predator Adults/Nymphs
Stethorus loi Predator Adults/Nymphs Taiwan Averrhoa carambola
Stethorus pauperculus Predator Adults/Eggs/Larvae/Nymphs
Typhlodromus pyri Predator Adults/Eggs/Larvae/Nymphs

References

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Anagnou-Veroniki M; Papaioannou-Souliotis P; Karanastasi E; Giannopolitis CN, 2008. New records of plant pests and weeds in Greece, 1990-2007. Hellenic Plant Protection Journal, 1(2):55-78. http://www.bpi.gr

Anupam Tagore, Putatunda, B. N., 2004. Mite pest of vegetable crops in Hisar (Haryana) India. Plant Protection Bulletin (Faridabad), 56(3/4), 32-36.

APPPC, 1987. Insect pests of economic significance affecting major crops of the countries in Asia and the Pacific region. Technical Document No. 135. Bangkok, Thailand: Regional Office for Asia and the Pacific region (RAPA).

Baker EW, 1983. Pests not known to occur in the United States or of limited distribution. No. 41: oriental red mite. Washington, USA: USDA-APHIS, 81-43.

Baker, E. W. , Pritchard, A. E. , 1960. The Tetranychoid mites of Africa. Hilgardia, 29(11), 455-574 pp.

Baker, E. W., 1975. Plant-feeding mites of Thailand (Tetranychidae, Tenuipalpidae, and Tuckerellidae). In: Plant Protection Service Technical Bulletin , (No.35) . 43pp.

CABI/EPPO, 1998. Distribution maps of quarantine pests for Europe (edited by Smith IM, Charles LMF). Wallingford, UK: CAB International, xviii + 768 pp.

CABI/EPPO, 2007. Eutetranychus orientalis. [Distribution map]. Distribution Maps of Plant Pests, No.June. Wallingford, UK: CABI, Map 688.

Chang DC; Leu TS, 1986. Seasonal population changes of spider mites on carambola and their chemical control. Plant Protection Bulletin, Taiwan, 28(3):263-272

Çobanoğlu, S., Can, M., 2014. Citrus brown mite; Eutetranychus orientalis (Klein 1936) (Acari: Tetranychidae), in Turkey. Akdeniz Üniversitesi Ziraat Fakültesi Dergisi, 27(1), 9-12. http://ziraatdergi.akdeniz.edu.tr/son-sayi

Deshpande RR; Shaw SS; Shrivastav RC; Mandloi KC, 1988. Relative toxicity of pesticides against red spider mite Eutetranychus orientalis (Klein) on Citrus. Pesticides, 22(3):45-46

Dhooria MS; Butani DK, 1984. Citrus mite, Eutetranychus orientalis (Klein) and its control. Pesticides, 18(10):35-38

Dhooria, M. S., Butani, D. K., 1982. Seasonal incidence of citrus mite, Eutetranychus orientalis and its predators. Indian Journal of Acarology, 7(2), 59-62.

Ehara, S., Wongsiri, T., 1975. The spider mites of Thailand (Acarina: Tetranychidae). Mushi, 48(13), 149-185.

El-Haidari, H. S., 1981. New records of mites and insects infesting date palms in the Near East and North Africa. Date Palm Journal, 1(1), 133-134.

El-Halawany ME; Ibrahim GA; Abdel-Samad MA, 1990. Mites inhabiting fig varieties. Agricultural Research Review, 68(1):39-48

EPPO, 1990. Specific quarantine requirements. EPPO Technical Documents, No. 1008. Paris, France: European and Mediterranean Plant Protection Organization.

EPPO, 2014. PQR database. Paris, France: European and Mediterranean Plant Protection Organization. http://www.eppo.int/DATABASES/pqr/pqr.htm

Gerson U; Kenneth R; Muttath TI, 1979. Hirsutella thompsonii, a fungal pathogen of mites. II. Host-pathogen interactions. Annals of Applied Biology, 91(1):29-40

González-Zamora, J. E., López, C., Avilla, C., 2011. Population studies of arthropods on Melia azedarach in Seville (Spain), with special reference to Eutetranychus orientalis (Acari: Tetranychidae) and its natural enemies. Experimental and Applied Acarology, 55(4), 389-400. doi: 10.1007/s10493-011-9476-y

Haq, M. A., Sumangala, K., 2003. Acarine regulators of water hyacinth in Kerala (India). Experimental and Applied Acarology, 29(1/2), 27-33. doi: 10.1023/A:1024203117619

Jeppson LR; Keifer HH; Baker EW, 1975. Mites Injurious to Economic Plants. Berkeley, USA: University of California Press.

Karuppuchamy, P., Mohanasundaram, M., 1987. New species and records of tetranychid mites (Tetranychidae: Acarina) from India. Entomon, 12(2), 89-94.

Khanjani, M., Khanjani, M., Seeman, O. D., 2017. New spider mites (Acari: Tetranychidae) of the genera Paraplonobia and Eurytetranychus from Iran, and a description of all life stages of Eutetranychus orientalis (Klein). Acarologia, 57(3), 465-491. http://www1.montpellier.inra.fr/CBGP/acarologia/export_pdf.php?id=4167&typefile=pdf

Khanjani, M., Ueckermann, E. A., 2008. A new species of Zetzellia Oudemans (Acari: Stigmaeidae) from west Iran. International Journal of Acarology, 34(3), 237-241.

Kumawat, K. C., Singh, S. P., 2002. Evaluation of insecticides and acaricides against Oriental mite infesting pomegranate. Annals of Plant Protection Sciences, 10(1), 137-139.

Massoudian, F., Khanjani, M., 2013. Mites associated with some medicinal plants (Asteraceae) in Hamedan, Iran. Journal of Crop Protection, 2(2), 209-218. https://www.semanticscholar.org/paper/MITES-ASSOCIATED-WITH-SOME-MEDICINAL-PLANTS-IN-IRAN-Masoudian-Khanjani/689ee1c5268d7d331365568dd33247382289e568#paper-header

Mazih A, 2015. Status of citrus IPM in the southern Mediterranean basin Morocco, North Africa. Acta Horticulturae [XII International Citrus Congress - International Society of Citriculture, Valencia, Spain.], No.1065:1097-1103. http://www.actahort.org/books/1065/1065_138.htm

McMurtry JA, 1985. Citrus. In: Helle W, Sabelis MW, eds. Spider Mites, their Biology, Natural Enemies and Control. World Crop Pests 1(B). Amsterdam, Netherlands: Elsevier, 339-347.

Meyer, M. K. P., 1987. African Tetranychidae (Acari: Prostigmata) - with reference to the world genera. In: Entomology Memoir, Department of Agriculture and Water Supply, Republic of South Africa , (No. 69) . iv + 175pp.

Nassar, O. A., Ghai, S., 1981. Taxonomic studies on tetranychoid mites infesting vegetable and fruit crops in Delhi and surrounding areas. 15(4), 333-396. doi: 10.1080/00305316.1981.10434337

Negm, M. W., 2014. Increasing knowledge of the mite fauna of the United Arab Emirates: new records and a checklist. Acarologia, 54(1), 113-120. doi: 10.1051/acarologia/20142118

Paz, Z., Gerson, U., Sztejnberg, A., 2007. Assaying three new fungi against citrus mites in the laboratory, and a field trial. BioControl, 52(6), 855-862. doi: 10.1007/s10526-006-9060-2

Pillai KS; Palaniswani MS, 1991. Integrated approach for the management of spider mite complex on cassava. Journal of Root Crops, 17:44-49.

PRITCHARD AE; BAKER EW, 1955. Mem. Pacif. Coast ent. Soc, 2. San Francisco, Cal., 7 + 472 pp.

Putatunda, B. N., Mathur, R. B., Mathur, S., 2002. Mites associated with some fruit trees in Hisar, Haryana, India. Indian Journal of Agricultural Research, 36(2), 88-95.

Rani, B. J., Sridhar, V., 2004. Record of arthropod pests on velvet bean, Mucuna pruriens var. utilis under Bangalore conditions. Journal of Medicinal and Aromatic Plant Sciences, 26(3), 505-506.

Rasmy, A. H., Zaher, M. A., Abou-Awad, B. A., 1972. Mites associated with deciduous fruit trees in U.A.R. Zeitschrift fur Angewandte Entomologie, 70(2), 179-183.

Sadana, G. L., Kanta, V., 1972. Susceptibility of different varieties of citrus to the citrus mite, Eutetranychus orientalis (Klein). Science and Culture, 38(12), 525-526.

Schotman CYL, 1989. Plant pests of quarantine importance to the Caribbean. RLAC-PROVEG, No. 21:80 pp.

Sharaf NS, 1989. Monitoring spider mite populations on lemon for effective control with flubenzimine and omethoate. Dirasat, 16(7):53-64.

Shun XuGen, Liu ZhenYu, Zhou ChengGang, Qiao LuQin, 1996. Damage of Eutetranychus orientalis to Euonymus japonicus. Plant Protection, 22(5), 29-30.

Smith IM; McNamara DG; Scott PR; Holderness M, 1997. Quarantine pests for Europe. Second Edition. Data sheets on quarantine pests for the European Union and for the European and Mediterranean Plant Protection Organization. Quarantine pests for Europe. Second Edition. Data sheets on quarantine pests for the European Union and for the European and Mediterranean Plant Protection Organization., Ed. 2:vii + 1425 pp.; many ref.

Smith IM; McNamara DG; Scott PR; Holderness M, 1997. Quarantine pests for Europe. Second Edition. Data sheets on quarantine pests for the European Union and for the European and Mediterranean Plant Protection Organization. Quarantine pests for Europe. Second Edition. Data sheets on quarantine pests for the European Union and for the European and Mediterranean Plant Protection Organization., Ed. 2:vii + 1425 pp.; many ref.

Smith-Meyer MKP, 1981. Mite pests of crops in southern Africa. Science Bulletin, Department of Agriculture and Fisheries, Republic of South Africa, (No. 397):65-67.

Smith-Meyer MKP, 1987. African Tetranychidae (Acari: Prostigmata) - with reference to the world fauna. Entomology Memoir, Department of Agriculture and Water Supply, Republic of South Africa, (No. 69):77-78, 80-82.

Soliman, Z. R., Al-Yousif, M. S., 1979. Prostigmatid mites of Saudi Arabia (Acari: Acariformes: Prostigmata). In: Zoological Society of Egypt Bulletin,29. 82-86.

Tanigoshi LK; Bahdousheh M; Babcock JM; Sawaqed R, 1990. Euseius scutalis (Athias-Henriot) a predator of Eutetranychus orientalis (Klein) (Acari: Phytoseiidae, Tetranychidae) in Jordan: Toxicity of some acaricides to E. orientalis. Arab Journal of Plant Protection, 8(2):120-114

Toroitich, F. J., Ueckermann, E. A., Theron, P. D., Knapp, M., 2009. The tetranychid mites (Acari: Tetranychidae) of Kenya and a redescription of the species Peltanobia erasmusi Meyer (Acari: Tetranychidae) based on males. Zootaxa, 2176, 33-47. doi: 10.5281/zenodo.189364

Walter, D. E., Halliday, R. B., Smith, D., 1995. The oriental red mite, Eutetranychus orientalis (Klein) (Acarina: Tetranychidae), in Australia. Journal of the Australian Entomological Society, 34(4), 307-308.

Waterhouse DF, 1993. The Major Arthropod Pests and Weeds of Agriculture in Southeast Asia. ACIAR Monograph No. 21. Canberra, Australia: Australian Centre for International Agricultural Research, 141 pp.

Zaher, M. A., Gomaa, E. A., El-Enany, M. A., 1982. Spider mites of Egypt (Acari: Tetranychidae). International Journal of Acarology, 8(2), 91-114.

Zriki, G., Shaabo, A., Boubou, A., 2015. A preliminary survey of the spider mites (Acari: Tetranychidae) in Latakia governorate of Syria. Acarologia, 55(3), 303-309. http://www1.montpellier.inra.fr/CBGP/acarologia/article.php?id=2173

Distribution References

Abad-Moyano R, Pina T, Dembilio Ó, Ferragut F, Urbaneja A, 2009. Survey of natural enemies of spider mites (Acari: Tetranychidae) in citrus orchards in eastern Spain. Experimental and Applied Acarology. 47 (1/2), 49-61. DOI:10.1007/s10493-008-9193-3

Abdel-Khalek A, Amer S, Momen F, 2011. Repellency and toxicity of extract from Francoeria crispa (Forsk.) to Eutetranychus orientalis (Klein) (Acari: Tetranychidae). Archives of Phytopathology and Plant Protection. 44 (5), 441-450. DOI:10.1080/03235400903093105

Alatawi F J, 2020. Field studies on occurrence, alternate hosts and mortality factors of date palm mite, Oligonychus afrasiaticus (McGregor) (Acari: Tetranychidae). Journal of the Saudi Society of Agricultural Sciences. 19 (2), 146-150. DOI:10.1016/j.jssas.2018.08.003

Anagnou-Veroniki M, Papaioannou-Souliotis P, Karanastasi E, Giannopolitis C N, 2008. New records of plant pests and weeds in Greece, 1990-2007. Hellenic Plant Protection Journal. 1 (2), 55-78. http://www.bpi.gr

Anupam Tagore, Putatunda B N, 2004. Mite pest of vegetable crops in Hisar (Haryana) India. Plant Protection Bulletin (Faridabad). 56 (3/4), 32-36.

APPPC, 1987. Insect pests of economic significance affecting major crops of the countries in Asia and the Pacific region. In: Technical Document No. 135, Bangkok, Thailand: Regional Office for Asia and the Pacific region (RAPA).

Atalla E A R, El-Atrouzy N, 1971. Survey of mites associated with vegetable crops in U.A.R. Agricultural Research Review. 49 (1), 116-117.

Baker E W , Pritchard A E, 1960. The Tetranychoid mites of Africa. Hilgardia. 29 (11), 455-574 pp.

Baker E W, 1975. Plant-feeding mites of Thailand (Tetranychidae, Tenuipalpidae, and Tuckerellidae). In: Plant Protection Service Technical Bulletin, 43pp.

Bhumannavar B S, Singh S P, Sulladmath V V, 1988. Evaluation of citrus germplasm for resistance to the oriental red mite, Eutetranychus orientalis (Klein) under tropical humid south Indian conditions. Tropical Pest Management. 34 (2), 193-198, 243, 247.

CABI, EPPO, 2007. Eutetranychus orientalis. [Distribution map]. In: Distribution Maps of Plant Pests, Wallingford, UK: CABI. Map 688. DOI:10.1079/DMPP/20073108412

CABI, Undated. Compendium record. Wallingford, UK: CABI

CABI, Undated a. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI

Chang D C, Leu T S, 1986. Seasonal population changes of spider mites on carambola and their chemical control. Plant Protection Bulletin, Taiwan. 28 (3), 263-272.

Çobanoğlu S, Can M, 2014. Citrus brown mite; Eutetranychus orientalis (Klein 1936) (Acari: Tetranychidae), in Turkey. Akdeniz Üniversitesi Ziraat Fakültesi Dergisi. 27 (1), 9-12. http://ziraatdergi.akdeniz.edu.tr/son-sayi

Debalina Mandal, Gupta S K, Sanyal A K, 2016. A report on new records of phytophagous mites on medicinal plants from Eastern Himalayan Region. Records of the Zoological Survey of India. 116 (4), 367-371.

Dhooria M S, 1983. Persistence of toxicity of pesticides applied to soil against eutetranychus orientalis on malta seedlings. Indian Journal of Acarology. 8 (1), 13-21. https://eurekamag.com/research/006/089/006089342.php

Dosse G, Musa S, 1967. Magon (Ser. sclent.). Tel-Amara. 2+] 23 pp.

Ehara S, 2004. A Collection of Spider Mites (Arachnida: Acari: Tetranychidae) from Sumatra. Species Diversity. 67-76. DOI:10.12782/specdiv.9.67

Elbadry E A, Afifi A M, Issa G I, Elbenhawy G M, 1968. Effectiveness of the predaceous mite Amblyseitts gossipi as a predator of three tetranychid mites (Acarina : Phytoseiidae). Zeitschrift für Angewandte Entomologie. 62 (43834), 189-194. DOI:10.1111/j.1439-0418.1968.tb04119.x

El-Haidari H S, 1981. New records of mites and insects infesting date palms in the Near East and North Africa. Date Palm Journal. 1 (1), 133-134.

El-Halawany M E, Ibrahim G A, Abdel-Samad M A, 1990. Mites inhabiting fig varieties. Agricultural Research Review. 68 (1), 39-48.

El-Kifl A H, Wahab A E A, Assem M A, Metwally A A, 1974. List of insects, mites and pests associated with leguminous crops in Egypt. Bulletin de la Societe Entomologique d'Egypte. 297-302.

EPPO, 2020. EPPO Global database. In: EPPO Global database, Paris, France: EPPO. https://gd.eppo.int/

Gerson U, Gafni A, Paz Z, Sztejnberg A, 2009. A tale of three acaropathogenic fungi in Israel: Hirsutella, Meira and Acaromyces. In: Experimental and Applied Acarology. 46 (43834) Netherlands: Springer. 183-194. DOI:10.1007/s10493-008-9202-6

González-Zamora J E, López C, Avilla C, 2011. Population studies of arthropods on Melia azedarach in Seville (Spain), with special reference to Eutetranychus orientalis (Acari: Tetranychidae) and its natural enemies. Experimental and Applied Acarology. 55 (4), 389-400. DOI:10.1007/s10493-011-9476-y

Haq M A, Sumangala K, 2003. Acarine regulators of water hyacinth in Kerala (India). Experimental and Applied Acarology. 29 (1/2), 27-33. DOI:10.1023/A:1024203117619

Ibrahim G A, El-Ghobashy M S, El-Sayed K M, Shoeib A A, 2005. Biological control of citrus brown mite Eutetranychus orientals using predatory mite, Neoseiulus californicus (McGregor) (Acari: Tetranychidae & Phytoseiidae) on citrus trees. Egyptian Journal of Agricultural Research. 83 (1), 131-139.

Karuppuchamy P, Mohanasundaram M, 1987. New species and records of tetranychid mites (Tetranychidae: Acarina) from India. Entomon. 12 (2), 89-94.

Khanjani M, Khanjani M, Seeman O D, 2017. New spider mites (Acari: Tetranychidae) of the genera Paraplonobia and Eurytetranychus from Iran, and a description of all life stages of Eutetranychus orientalis (Klein). Acarologia. 57 (3), 465-491. http://www1.montpellier.inra.fr/CBGP/acarologia/export_pdf.php?id=4167&typefile=pdf

Khanjani M, Ueckermann E A, 2008. A new species of Zetzellia Oudemans (Acari: Stigmaeidae) from west Iran. International Journal of Acarology. 34 (3), 237-241.

Knorr L C, Moin Shah S, 1971. World Citrus problems-V. Nepal. Plant Protection Bulletin, F.A.O. 19 (4), 73-79.

Kumawat K C, Singh S P, 2002. Evaluation of insecticides and acaricides against Oriental mite infesting pomegranate. Annals of Plant Protection Sciences. 10 (1), 137-139.

Ledesma C, Wong M E, Vela J M, Jacas J A, Boyero J R, 2011. Population dynamics of the citrus oriental mite, Eutetranychus orientalis (Klein) (Acari: Tetranychidae), and its mite predatory complex in southern Spain. IOBC/WPRS Bulletin. 83-92. http://www.iobc-wprs.org/pub/bulletins/bulletin_2011_62_table_of_contents_abstracts.pdf

Marić I, Međo I, Marčić D, Petanović R, Jovanović S, Ueckermann EA, 2021. Spider mites (Acari: Tetranychidae) from Serbia: New species for the country and the Balkan Peninsula, with a key to all known Serbian species. Systematic and Applied Acarology. 26 (1), 304-316.

Massoudian F, Khanjani M, 2013. Mites associated with some medicinal plants (Asteraceae) in Hamedan, Iran. Journal of Crop Protection. 2 (2), 209-218. https://www.semanticscholar.org/paper/MITES-ASSOCIATED-WITH-SOME-MEDICINAL-PLANTS-IN-IRAN-Masoudian-Khanjani/689ee1c5268d7d331365568dd33247382289e568#paper-header

Mazih A, 2015. Status of citrus IPM in the southern Mediterranean basin Morocco, North Africa. Acta Horticulturae. 1097-1103. http://www.actahort.org/books/1065/1065_138.htm

Meyer M K P, 1987. African Tetranychidae (Acari: Prostigmata) - with reference to the world genera. In: Entomology Memoir, Department of Agriculture and Water Supply, Republic of South Africa, iv + 175pp.

Nassar O A, Ghai S, 1981. Taxonomic studies on tetranychoid mites infesting vegetable and fruit crops in Delhi and surrounding areas. 15 (4), 333-396. DOI:10.1080/00305316.1981.10434337

Negm M W, 2014. Increasing knowledge of the mite fauna of the United Arab Emirates: new records and a checklist. Acarologia. 54 (1), 113-120. DOI:10.1051/acarologia/20142118

NPPO of the Netherlands, 2013. Pest status of harmful organisms in the Netherlands., Wageningen, Netherlands:

Osman A A, Rasmy A H, Undated. The role of alfalfa in dispersing tetranychid mites to other crops. Bulletin of the Entomological Society of Egypt. 279-283.

Paz Z, Gerson U, Sztejnberg A, 2007. Assaying three new fungi against citrus mites in the laboratory, and a field trial. BioControl. 52 (6), 855-862. DOI:10.1007/s10526-006-9060-2

Putatunda B N, Mathur R B, Mathur S, 2002. Mites associated with some fruit trees in Hisar, Haryana, India. Indian Journal of Agricultural Research. 36 (2), 88-95.

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