Chloris barbata (purpletop chloris)

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Inflorescence and leaves - colour illustration

A tufted, erect, annual or short-lived perennial grass, 0.3-1.0 m or more tall.

NOVARTIS

Inflorescence - line drawing

Inflorescence terminal, composed of a whorl of 5-15 digitate spikes which are densely clustered.

NOVARTIS

Spikelets - line drawing

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Leaf - line drawing

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Identity

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Preferred Scientific Name

Chloris barbata Sw., 1797

Preferred Common Name

purpletop chloris

Other Scientific Names

Andropogon barbatus L., 1771, non L., 1759
Chloris inflata Link, 1821
Chloris longifolia Steud., 1854
Chloris paraguayensis Steud., 1854
Chloris refuscens Steud., 1854, non Lag., 1805

International Common Names

English: peacock plumegrass; plush grass; purple top; swollen fingergrass
Spanish: zacate borrego

Local Common Names

Cuba: pata de gallina
Indonesia: rumput jejarongan; suket cakar ayam
Japan: murasaki-higeshiba
Malaysia: kilen
Philippines: banuko; korokorosan
Sri Lanka: kondai pul; mayuru tana
Thailand: yaa rangnok
USA/Hawaii: mau'u lei

EPPO code

CHRBA (Chloris barbata)

Taxonomic Tree

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Domain: Eukaryota
Kingdom: Plantae
Phylum: Spermatophyta
Subphylum: Angiospermae
Class: Monocotyledonae
Order: Cyperales
Family: Poaceae
Genus: Chloris
Species: Chloris barbata

Notes on Taxonomy and Nomenclature

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C. barbata belongs to the tribe Chlorideae (Poaceae).

C. barbata is based on Andropogon barbata which is an illegitimate name, being a later homonym of A. barbatus, the names being based on different plants. The epithet barbata is not, however, illegitimate in the genus Chloris and Swartz was perfectly in order in calling his plant C. barbata, there being no other epithet available. As the citation C. barbata is ambiguous, the Code permits us to forget that Linnaeus made the illegitimate combination. Thus, C. barbata is regarded as a new name dating from 1797 and should be cited without acknowledgement to Linnaeus (Bor, 1960; Cope, 1982). Chloris inflata, a name often applied to this species, is predated by C. barbata.

Description

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C. barbata is a tufted, erect, annual or short-lived perennial grass. It is 0.3-1.0 m or more tall, largely glabrous, with a short life span, heading and flowering all year round. The erect and branching stems, which are sometimes bent at the base, are smooth and usually flattened. They are purple or pink at the base, simple or branched, 3-5-noded, rooting at the lower nodes.

Leaves

The leaf blades are flat and narrow, linear-lanceolate, 10-20 cm long (the upper ones shorter), 2-3 mm wide and usually bluish-green with rough edges. They often have long, scattered hairs on the upper surface, near the base. The sheaths are smooth and 2-6 cm long; usually less than half as long as the internodes. They are compressed, keeled and closely overlapping, with glabrous or bearded orifice. The ligule is 0.5-1.0 mm long, membraneous and fringed with short hairs.

Inflorescence

Terminal, composed of a whorl of 5-15 digitate spikes which are densely clustered. The spikes are usually ascending, purple and 5-8 cm long with three-flowered spikelets (one fertile flower). These are purplish and densely overlapping, with three slender awns. The glumes are unequal, narrow, acute and membraneous except for a single green nerve. The first is 1-1.5 mm long and the second 2-2.5 mm long. There are almost always three florets which are often purple. The rachilla internode is 1 mm long. The lemmas are three-nerved. The first lemma is obovate, keeled and 2-2.5 mm long. It is sparsely to densely pilose on the margins and the keel. The awn is 5-10 mm long. The palea is 2-2.5 mm long and nearly as broad as the lemma with marginal keels. The apical rudiment is approximately 1 mm long and consists of two inflated, triangular-truncate, thin, glabrous, sterile lemmas, one within the other, each with an awn 3-5 mm long.

Chromosome number: 2n = 20.

Grain

C. barbata grain is pale brown, tapering at both ends and 1-2 mm long, enclosed within the persistent lemma and palea.

C. barbata is propagated by seed.

Distribution

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The origin of C. barbata is uncertain. Bor (1960) states that it is distributed through the "tropics of South-East Asia, introduced elsewhere (but considered to be native in Tropical America". Other floras simply accept that it is now widely distributed in tropical and some subtropical regions of all continents, with the possible exception of southern Africa.

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Last updated: 25 Feb 2021

Continent/Country/Region	Distribution	Origin	Invasive	Reference
Africa
Benin	Present			Holm et al. (1979)
Côte d'Ivoire	Present			Clayton (1972)
Egypt	Present			Holm et al. (1979)
Ghana	Present			Holm et al. (1979)
Kenya	Present			Holm et al. (1979)
Madagascar	Present			Bosser (1969)
Mauritius	Present			Holm et al. (1979)
Nigeria	Present			Holm et al. (1979)
Sudan	Present			Holm et al. (1979)
Tanzania	Present			Holm et al. (1979)
-Zanzibar Island	Present			Clayton et al. (1974)
Asia
Cambodia	Present			Holm et al. (1979)
China	Present			Holm et al. (1979)
Hong Kong	Present			Holm et al. (1979)
India	Present			Holm et al. (1979) Munirathnam and Kumar (2014) Devi et al. (2015) Nayak and Satapathy (2015)
-Andhra Pradesh	Present			Munirathnam and Kumar (2014)
-Odisha	Present			Nayak and Satapathy (2015)
-Tamil Nadu	Present			Devi et al. (2015)
Indonesia	Present			Holm et al. (1979)
Iran	Present			Holm et al. (1979)
Malaysia	Present			Holm et al. (1979)
North Korea	Present			Holm et al. (1979)
Pakistan	Present			Cope (1982)
Philippines	Present			Holm et al. (1979)
Sri Lanka	Present			Clayton et al. (1994)
Taiwan	Present			Anon (1978)
Thailand	Present			Holm et al. (1979)
Vietnam	Present			Holm et al. (1979)
North America
Cuba	Present	Introduced	Invasive	Oviedo Prieto et al. (2012)
Jamaica	Present			Holm et al. (1979)
Mexico	Present			Hafliger and Scholz (1981)
Puerto Rico	Present			Holm et al. (1979)
United States	Present			Holm et al. (1979)
-Hawaii	Present			Holm et al. (1979)
-Louisiana	Present			Macgregor and Allen (1990)
Oceania
Australia	Present			Holm et al. (1979) Blanche et al. (2003) Grice et al. (2004)
-Queensland	Present			Stanley and Ross (1989) Grice et al. (2004)
-Western Australia	Present			Blanche et al. (2003)
Fiji	Present			Holm et al. (1979)
French Polynesia	Present			Whistler (1994)
Guam	Present			Whistler (1994)
Marshall Islands	Present			Taylor (1950)
Palau	Present			Whistler (1994)
Papua New Guinea	Present			Henty (1969)
Samoa	Present			Sauerborn and Sauerborn (1984) Whistler (1994)
Tonga	Present			Whistler (1994)
South America
Argentina	Present			Hafliger and Scholz (1981)
Bolivia	Present			Hafliger and Scholz (1981)
Brazil	Present			Hafliger and Scholz (1981)
Colombia	Present			Hafliger and Scholz (1981)
Peru	Present			Hafliger and Scholz (1981)
Venezuela	Present			Hafliger and Scholz (1981)

Habitat

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C. barbata is possibly native to tropical America but is now widespread at low altitudes throughout the tropics. It prefers dry conditions and is common in coastal areas. It is a common weed in Mexico, South America, the West Indies, Papua New Guinea and Sri Lanka. In the Hawaiian islands, it is a common species in the arid coastal zone (Sohmer and Gustafson, 1987). It is found in dryland field crops, pastures, sugarcane, wastelands, abandoned cultivation, railway embankments, roadsides, borders of plantation crops and on levees in lowland rice fields.

In South-East Asia, it occurs in lowland ricefields, lawns, by roadsides and railway tracks and similar disturbed situations. It tends to be saline tolerant and is common in desert pans and littoral areas and on the fringes of salt meadows and mangrove swamps (Lazarides, 1980).

Host Plants and Other Plants Affected

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Plant name	Family	Context
Arachis hypogaea (groundnut)	Fabaceae	Main
Carica papaya (pawpaw)	Caricaceae	Main
Cocos nucifera (coconut)	Arecaceae	Other
Coffea arabica (arabica coffee)	Rubiaceae	Main
Macadamia ternifolia (Queensland nut)	Proteaceae	Main
Nicotiana tabacum (tobacco)	Solanaceae	Main
Oryza sativa (rice)	Poaceae	Main
Saccharum officinarum (sugarcane)	Poaceae	Main

Biology and Ecology

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C. barbata appears to vary considerably in its habit in different parts of the world, being variously described as annual or perennial and erect or stoloniferous and rooting at the nodes. It presumably propagates mainly by seed, but information on germination biology is not available. As noted under Habitat, it is a plant of relatively dry conditions, often near the coast and apparently tolerant of saline conditions.

C. barbata is a host of a number of rice insect pests and diseases, including the white-backed planthoppers, Sogatella furcifera (Vaidya and Kalode, 1982; Catindig et al., 1988, 1991) and Sogatodes pusanus [Tagosodes pusanus] (Catindig et al., 1989), the rice bug Leptocorisa oratorius (Rajapakse and Kulasekera, 1980), the rice earcutting caterpillar Mythimna separata (Catindig et al., 1991, 1994), the rice-feeding tiger moth Creatonotus gangis (Catindig et al., 1991, 1993), the cereal thrips Haplothrips ganglbaurei (Ananthakrishnan and Thangavelu, 1976), the rice whitefly Aleurocybotus indicus (Alam, 1989), and sheath blight Rhizoctonia solani [Thanatephorus cucumeris] (Kannaiyan and Prasad, 1979). It is also an ovipositional host of the rice leaffolder, Cnaphalocrocis medinalis (Barrion et al., 1991).

It is also the principal alternative host of the grass seed-feeding thrips, Chirothrips mexicanus, which infests the cultivated crop pearl millet, Pennisetum typhoides [Pennisetum glaucum] (Ananthakrishnan and Thirumalai, 1977).

Impact

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C. barbata is reported as a serious weed in Australia, Korea and Thailand, a principal weed in Cambodia and India, and a common weed in Hawaii, Malaysia and the Sudan (Holm et al., 1979). It was a serious weed of sugarcane in Hawaii before trifluralin was used for control (Santo, personal communication).

C. barbata is a common weed in sugarcane, tree crops (papayas, macadamia nuts, coffee) and lawns in Hawaii (R.K. Nishomoto, University of Hawaii, personal communication) and occurs in groundnuts in India (Rajan et al., 1981) and tobacco in the Philippines (Pancho and Obien, 1983) and Thailand (Suwanarak et al., 1986). It occurs infrequently in coconut plantations in the northern Marshall Islands (Taylor, 1950).

It has been reported as a weed of upland rice in India, Indonesia, Philippines, Sri Lanka, Thailand and Vietnam, of dry-seeded rice in India and Thailand, of wet-seeded rice (sprouted seeds sown on puddled soil) in Thailand, and of transplanted rice in India and the Philippines. It has also been reported as occurring in rice seedling nurseries in Thailand and in lowland rice in Indonesia (Moody, 1989).

Yield losses of up to 20% in sugarcane are likely with heavy infestations of C. barbata. On Oahu, Hawaii, C. barbata developed resistance to herbicides and eventually entire fields were infested with high populations of C. barbata up to 1.5 m tall. In some cases, where sugarcane growth was poor, infestations were so severe that the crop was ploughed under and the field replanted (LT Santo, Hawaiian Sugar Planters' Association, personal communication).

Fresh root extracts of C. barbata collected from South Arcot, Tamil Nadu, inhibited growth of ragi (Eleusine coracana) seeds (Jeyamurthy and Lakshmanachary, 1989).

C. barbata is grazed by stock when young but it soon becomes unpalatable and, thereafter, it is a pest (Whitney et al., 1939). According to Henty (1969), it is useless for grazing. The purplish, feathery-looking spikes are used in hat leis in Hawaii (Neal, 1965).

Similarities to Other Species/Conditions

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C. barbata is recognized by a combination of characteristics, such as its loosely appressed spikes, three flowers, three-awned spikelet which is often purple, almost entire lemmas with subequal awns and the inflated truncate lemmas of the imperfect florets. The lowest (fertile) floret is distinguished by its long-bearded callus and submarginal fringe of stiff white hairs on the upper part of the lemma (Clayton et al., 1994).

A similar species, C. virgata (feather fingergrass or feathertop Rhodes grass), differs in having white to yellowish-brown spikes and only two (rather than three) distinct awns (Whistler, 1994). C. gayana (Rhodes grass), a valuable fodder grass, is superficially similar but the spikes are yellow-brown rather than purple and the awns are much shorter, less than 1.5 times as long as the lemmas (Kleinschmidt and Johnson, 1977). Another weedy species, C. pilosa also has very short awns. C. prieurii has long awns, but there are 4-6 per spikelet. The annual C. pycnothrix has very distinctive short, blunt leaves.

Prevention and Control

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Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.

Cultural Control

Good cultural practices, including good tillage, use of vigorous cultivars and timing of water and fertilizer applications maximize sugarcane growth and canopy closure (Santo, 1989). Healthy growing sugarcane will grow above C. barbata and shade it out and yield losses will be slight, if any (LT Santo, Hawaiian Sugar Planters' Association, personal communication).

Chemical Control

In Hawaii, pre-emergence herbicides are the primary method of controlling weeds in sugarcane, with post-emergence treatments as a back-up. Grass weeds, including C. barbata, are difficult to control with post-emergence herbicides without damaging the crop. Ametryn or diuron + atrazine account for 80% of the herbicides used. A pre-emergence application in irrigated fields gives control for 5-8 weeks. Terbacil or hexazinone may be added to control C. barbata. Metsulfuron also gives good control of C. barbata. Ametryn or diuron, atrazine and 2,4-DB are used with dalapon or hexazinone for perennial grasses. Glyphosate is spot-applied to tall grass stands, with 2,4-D if broadleaved weeds are present (Santo, 1989). C. barbata developed resistance to ametryn and diuron. However, it is susceptible to pre-emergence applications of trifluralin or pendimethalin and is no longer a problem in sugarcane fields in Hawaii (LT Santo, Hawaiian Sugar Planters' Association, personal communication). Barnes and Chandapillai (1972) suggest the use of methylarsonic acid plus 2,4-D.

Glyphosate is used to control C. barbata in tree crops in Hawaii and spot applications of glyphosate or glufosinate are used in lawns (RK Nishimoto, University of Hawaii, personal communication).

In Andhra Pradesh, India, fluchloralin effectively controlled C. barbata and other narrow-leaved weeds in irrigated groundnuts (Rajan et al., 1981).

In tobacco in Thailand, clopomydim, fenoxaprop-ethyl, haloxyfop-methyl and fluazifop-butyl gave good control of the narrow-leaved weeds, including C. barbata, but could not control all of the broadleaved weeds. Fluazifop-butyl + bifenox, haloxyfop-methyl + dimethazone [withdrawn] and fenoxaprop-ethyl + dimethazone [withdrawn] controlled all the annual weeds for longer than 4 weeks and did not damage the crop (Suwanarak et al., 1986).

References

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Alam MS, 1989. Whitefly (Hemiptera: Aleyrodidae) - a potential pest of rice in West Africa. International Rice Research Newsletter, 14(3):38-39

Ananthakrishnan TN; Thangavelu K, 1976. The cereal thrips Haplothrips ganglbaueri Schmutz with particular reference to the trends of infestation on Oryza sativa and the weed Echinochloa crusgalli. Proceedings of the Indian Academy of Sciences, B, 83(5):196-201

Ananthakrishnan TN; Thirumalai G, 1977. The grass seed infesting thrips Chirothrips mexicanus Crawford on Pennisetum typhoides and its principal alternate host, Chloris barbata. Current Science, 46(6):193-194

Anon, 1978. Flora of Taiwan, Volume 5. Taipei, China: Epoch Publishing.

Barnes DE; Chandapillai MM, 1972. Common Malaysian Weeds and their Control. Kuala Lumpur, Malaysia: Ansul (Malaysia) Sdn. Berhad.

Bor NL, 1960. The Grasses of Burma, Ceylon, India and Pakistan (Excluding Bambusae). Oxford, UK: Pergamon Press.

Bosser J, 1969. Graminees des paturages et des cultures a Madagascar. Paris, France: ORSTOM.

Catindig JLA; Barrion AT; Litsinger JA, 1988. Host range of yellow rice borer, brown and whitebacked planthoppers. Philippine Association of Entomologists, Abstracts of Papers, 19th Anniversary and Annual Convention, Pest Control Council of the Philippines, 3-7 May 1988, Cebu City, Philippines.

Catindig JLA; Barrion AT; Litsinger JA, 1989. Life history and hosts of Sogatodes pusanus (Distant) (Hemiptera: Delphacidae). International Rice Research Newsletter, 19(1):23-24.

Catindig JLA; Barrion AT; Litsinger JA, 1991. Evaluation of weed alternate hosts to selected rice insect pests. Paper presented at the 22nd Annual Convention of the Pest Management Council of the Philippines, 8-10 May 1991, Manila, Philippines.

Catindig JLA; Barrion AT; Litsinger JA, 1993. Developmental biology and host plant range of rice-feeding tiger moth Creatonotus gangis (L.). International Rice Research Notes, 18(3):34-35

Catindig JLA; Barrion AT; Litsinger JA, 1994. Development biology and host plant range of rice ear-cutting caterpillar Mythimna separata (Walker). International Rice Research Notes, 19(1):23-24.

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Clayton WD; Davidse G; Gould F; Lazarides M; Soderstrom TR, 1994. A Revised Handbook to the Flora of Ceylon. Volume VIII. Poaceae. New Delhi, India: Amerind.

Cope TA, 1982. 143. Poaceae. In: Nasir E, Ali SI, eds. Flora of Pakistan. Karachi, Pakistan: University of Karachi.

Hafliger E; Scholz H, 1981. Grass weeds 2. Grass weeds 2. Ciba-Geigy Ltd. Basle Switzerland, xx + 137 + 23 pp.

Haselwood EL; Motter GG, 1966. Handbook of Hawaiian weeds. Hawaii, USA: Hawaiian Sugar Planters' Association, 479pp.

Henty EE, 1969. A Manual of the Grasses of New Guinea. Botany Bulletin No. 1, Lae, Papua New Guinea: Division of Botany.

Hitchcock AS, 1922. The Grasses of Hawaii. Memoirs of the Bernice Pauahi Bishop Museum, Volume VIII, Number 3. Honolulu, Hawaii, USA: Bishop Museum.

Holm LG; Pancho JV; Herberger JP; Plucknett DL, 1979. A geographical atlas of world weeds. New York, USA: John Wiley and Sons, 391 pp.

Jeyamurthy A; Lakshmanachary AS, 1989. Effect of root of Chloris barbata Sw. and Digitaria griffithii Stapf. on the germination of ragi seeds. Indian Journal of Applied and Pure Biology, 4(2):123-125

Kannaiyan S; Prasad NN, 1979. Sheath blight incidence in weed hosts. International Rice Research Newsletter, 4(3):17

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Lazarides M, 1980. The Tropical Grasses of Southeast Asia. Vaduz, Germany: Strauss and Cramer, 225 pp.

Macgregor JR; Allen CM, 1990. Chloris inflata (Poaceae), new to Louisiana. SIDA, Contributions to Botany, 14(2):313.

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Noda K; Teerawatsakul M; Praknogvongs C; Chaiwirtnukul L, 1984. Major Weeds in Thailand. Project Manual No. 1. Bangkok, Thailand: National Weed Science Research Institute.

Oviedo Prieto R; Herrera Oliver P; Caluff MG, et al. , 2012. National list of invasive and potentially invasive plants in the Republic of Cuba - 2011. (Lista nacional de especies de plantas invasoras y potencialmente invasoras en la República de Cuba - 2011). Bissea: Boletín sobre Conservación de Plantas del Jardín Botánico Nacional de Cuba, 6(Special Issue 1):22-96.

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Distribution References

Anon, 1978. Flora of Taiwan., 5 Taipei, China: Epoch Publishing.

Blanche K R, Tran-Nguyen L T T, Gibb K S, 2003. Detection, identification and significance of phytoplasmas in grasses in northern Australia. Plant Pathology. 52 (4), 505-512. DOI:10.1046/j.1365-3059.2003.00871.x

Bosser J, 1969. (Graminees des paturages et des cultures a Madagascar)., Paris, France: ORSTOM.

CABI, Undated. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI

Clayton W D, Phillips S M, Renvoize S A, 1974. Flora of Tropical East Africa. Gramineae (Part 2). In: Flora of Tropical East Africa. Gramineae (Part 2). London, UK: Crown Agents for Oversea Governments and Administrations. 274pp.

Clayton WD, 1972. 202. Gramineae. In: Flora of West Tropical Africa, [ed. by Hutchinson J, Dalziel JM, Hepper FN]. London, UK: Crown Agents. 349-512.

Clayton WD, Davidse G, Gould F, Lazarides M, Soderstrom TR, 1994. A Revised Handbook to the Flora of Ceylon. Volume VIII. Poaceae., New Delhi, India: Amerind.

Cope TA, 1982. 143. Poaceae. In: Flora of Pakistan, [ed. by Nasir E, Ali SI]. Karachi, Pakistan: University of Karachi.

Devi M R, Madhavan S, Baskaran A, Thangaratham T, 2015. Ethno medicinal aspects of weeds from paddy field in Thiruvarur district, Tamil Nadu, India. World Journal of Pharmaceutical Research. 4 (11), 1909-1920. http://www.wjpr.net/dashboard/abstract_id/4153

Grice A C, Lawes R A, Abbott B N, Nicholas D M, Whiteman L V, 2004. How abundant and widespread are riparian weeds in the dry tropics of north-east Queensland? In: Weed management: balancing people, planet, profit. 14th Australian Weeds Conference, Wagga Wagga, New South Wales, Australia, 6-9 September 2004: papers and proceedings. [ed. by Sindel B M, Johnson S B]. Sydney, Australia: Weed Society of New South Wales. 173-175.

Hafliger E, Scholz H, 1981. Grass weeds 2. In: Grass weeds 2. Basle, Switzerland: Ciba-Geigy Ltd. xx + 137 + 23 pp.

Henty EE, 1969. A Manual of the Grasses of New Guinea. In: Botany Bulletin No. 1, Lae, Papua New Guinea: Division of Botany.

Holm L, Pancho J V, Herberger J P, Plucknett D L, 1979. A geographical atlas of world weeds. New York, Chichester (), Brisbane, Toronto, UK: John Wiley and Sons. xlix + 391 pp.

Macgregor JR, Allen CM, 1990. Chloris inflata (Poaceae), new to Louisiana. In: SIDA, Contributions to Botany, 14 (2) 313.

Munirathnam P, Kumar K A, 2014. Survey of weed flora in major crops of Nandyal region in Kurnool district of Andhra Pradesh. Journal of Research ANGRAU. 42 (3/4), 142-146. http://www.angrau.ac.in/journalofresearchangrau.php

Nayak S K, Satapathy K B, 2015. Diversity, uses and origin of invasive alien plants in Dhenkanal district of Odisha, India. International Research Journal of Biological Sciences. 4 (2), 21-27. http://www.isca.in/IJBS/Archive/v4/i2/4.ISCA-IRJBS-2014-223.pdf

Oviedo Prieto R, Herrera Oliver P, Caluff M G, et al, 2012. National list of invasive and potentially invasive plants in the Republic of Cuba - 2011. (Lista nacional de especies de plantas invasoras y potencialmente invasoras en la República de Cuba - 2011). Bissea: Boletín sobre Conservación de Plantas del Jardín Botánico Nacional de Cuba. 6 (Special Issue No. 1), 22-96.

Sauerborn E, Sauerborn J, 1984. Plants of cropland in Western Samoa with special reference to taro. Plits. 2 (4), 331pp.

Stanley T D, Ross E M, 1989. Flora of south-eastern Queensland. Volume 3. Brisbane, Australia: Queensland Department of Primary Industries. 532 pp.