Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide

Datasheet

Ceroplastes floridensis
(soft scale)

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Datasheet

Ceroplastes floridensis (soft scale)

Summary

  • Last modified
  • 27 March 2018
  • Datasheet Type(s)
  • Pest
  • Natural Enemy
  • Host Animal
  • Preferred Scientific Name
  • Ceroplastes floridensis
  • Preferred Common Name
  • soft scale
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Metazoa
  •     Phylum: Arthropoda
  •       Subphylum: Uniramia
  •         Class: Insecta

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Pictures

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PictureTitleCaptionCopyright
Florida wax scales on citrus leaf.
TitleWax scales
CaptionFlorida wax scales on citrus leaf.
CopyrightNOVARTIS Crop Protection AG, Basel Switzerland
Florida wax scales on citrus leaf.
Wax scalesFlorida wax scales on citrus leaf.NOVARTIS Crop Protection AG, Basel Switzerland

Identity

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Preferred Scientific Name

  • Ceroplastes floridensis Comstock, 1881

Preferred Common Name

  • soft scale

Other Scientific Names

  • Cerostegia floridensis (Comstock) De Lotto, 1969
  • Paracerostegia floridensis (Comstock) Tang, 1991

International Common Names

  • English: Florida wax scale; wax scale, Florida
  • Spanish: cochinilla perla pociedrica; escama cerosa de la Florida; escama de cera; guagua de cera de Florida
  • French: ceroplaste de Floride; cochenille de Floride

Local Common Names

  • Germany: Florida schildlaus; Schildlaus, Florida-; Wachsschildlaus, Florida-
  • Israel: knimat hashaava hafloridit
  • Italy: Ceroplaste di Florida
  • Turkey: kanli balsira

EPPO code

  • CERPFL (Ceroplastes floridensis)

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta
  •                     Order: Hemiptera
  •                         Suborder: Sternorrhyncha
  •                             Unknown: Coccoidea
  •                                 Family: Coccidae
  •                                     Genus: Ceroplastes
  •                                         Species: Ceroplastes floridensis

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

Continent/Country/RegionDistributionLast ReportedOriginFirst ReportedInvasiveReferenceNotes

Asia

BangladeshPresentCABI/EPPO, 2010; EPPO, 2014
Brunei DarussalamPresentCABI/EPPO, 2010; EPPO, 2014
ChinaPresentCABI/EPPO, 2010; EPPO, 2014
-FujianPresentCABI/EPPO, 2010; EPPO, 2014
-GuizhouPresentCABI/EPPO, 2010; EPPO, 2014
-HebeiPresentCABI/EPPO, 2010; EPPO, 2014
-Hong KongPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-HunanPresentCABI/EPPO, 2010; EPPO, 2014
-JiangsuPresentCABI/EPPO, 2010; EPPO, 2014
-MacauPresentCABI/EPPO, 2010; EPPO, 2014
-YunnanPresentCABI/EPPO, 2010; EPPO, 2014
-ZhejiangPresentCABI/EPPO, 2010; EPPO, 2014
Georgia (Republic of)PresentCABI/EPPO, 2010; EPPO, 2014
IndiaPresentCABI/EPPO, 2010; EPPO, 2014
-Andaman and Nicobar IslandsPresentNHM, 1991; CABI/EPPO, 2010; EPPO, 2014
-AssamPresentCABI/EPPO, 2010; EPPO, 2014
-BiharPresentCABI/EPPO, 2010; EPPO, 2014
-KarnatakaPresentCABI/EPPO, 2010; EPPO, 2014
-KeralaPresentCABI/EPPO, 2010; EPPO, 2014
-MaharashtraPresentNHM, 1991; CABI/EPPO, 2010; EPPO, 2014
-OdishaPresentNHM, 1982; CABI/EPPO, 2010; EPPO, 2014
-Tamil NaduPresentCABI/EPPO, 2010; EPPO, 2014
-Uttar PradeshPresentCABI/EPPO, 2010; EPPO, 2014
-West BengalPresentCABI/EPPO, 2010; EPPO, 2014
IndonesiaPresentCABI/EPPO, 2010; EPPO, 2014
-Irian JayaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-JavaPresentNHM, pre-1940; CABI/EPPO, 2010; EPPO, 2014
-SulawesiPresentNHM, 1985; CABI/EPPO, 2010; EPPO, 2014
IranPresentCABI/EPPO, 2010; EPPO, 2014
IsraelPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
JapanPresentCABI/EPPO, 2010; EPPO, 2014
-Bonin IslandPresentCABI/EPPO, 2010
-HonshuPresentCABI/EPPO, 2010; EPPO, 2014
-KyushuPresentCABI/EPPO, 2010; EPPO, 2014
-Ryukyu ArchipelagoPresentCABI/EPPO, 2010; EPPO, 2014
-ShikokuPresentCABI/EPPO, 2010; EPPO, 2014
JordanPresentCABI/EPPO, 2010; EPPO, 2014
Korea, Republic ofPresentCABI/EPPO, 2010; EPPO, 2014
LebanonPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
MalaysiaPresentCABI/EPPO, 2010; EPPO, 2014
-Peninsular MalaysiaPresentCABI/EPPO, 2010; EPPO, 2014
-SabahPresentCABI/EPPO, 2010; EPPO, 2014
-SarawakPresentCABI/EPPO, 2010; EPPO, 2014
OmanPresentNHM, 1987; CABI/EPPO, 2010; EPPO, 2014
PakistanPresentCABI/EPPO, 2010; EPPO, 2014
Saudi ArabiaPresentNHM, 1989; CABI/EPPO, 2010; EPPO, 2014
Sri LankaPresentCABI/EPPO, 2010; EPPO, 2014
SyriaPresentCABI/EPPO, 2010; EPPO, 2014
TaiwanPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
TurkeyPresentBen-Dov, 1993; CABI/EPPO, 2010; Ülgentürk et al., 2012; EPPO, 2014
VietnamPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014

Africa

ComorosPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
EgyptPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
KenyaPresentNHM, 1929; CABI/EPPO, 2010; EPPO, 2014
LibyaPresentCABI/EPPO, 2010; EPPO, 2014
MadagascarPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
MauritiusPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
MozambiquePresentCABI/EPPO, 2010; EPPO, 2014
RéunionPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
SeychellesPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
Sierra LeonePresentCABI/EPPO, 2010; EPPO, 2014
South AfricaPresentNHM, 1954; CABI/EPPO, 2010; EPPO, 2014
TanzaniaPresentCABI/EPPO, 2010; EPPO, 2014
-ZanzibarPresentNHM, 1913
TunisiaPresentJendoubi et al., 2012
UgandaPresentCIE, 1982; CABI/EPPO, 2010; EPPO, 2014

North America

BermudaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
MexicoPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
USAPresentCABI/EPPO, 2010; EPPO, 2014
-AlabamaPresentCABI/EPPO, 2010; EPPO, 2014
-CaliforniaAbsent, unreliable recordCABI/EPPO, 2010
-FloridaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-GeorgiaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-HawaiiPresentCABI/EPPO, 2010; EPPO, 2014
-LouisianaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-MarylandPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-MississippiPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-MissouriPresentCABI/EPPO, 2010; EPPO, 2014
-New JerseyPresentCABI/EPPO, 2010; EPPO, 2014
-New MexicoPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-New YorkPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-North CarolinaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-South CarolinaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-TennesseePresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-TexasPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-VirginiaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-WashingtonPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014

Central America and Caribbean

Antigua and BarbudaPresentCABI/EPPO, 2010; EPPO, 2014
BahamasPresentCABI/EPPO, 2010; EPPO, 2014
BarbadosPresentCABI/EPPO, 2010; EPPO, 2014
BelizePresentEPPO, 2014
British Virgin IslandsPresentCABI/EPPO, 2010; EPPO, 2014
Cayman IslandsPresentNHM, 1970; CABI/EPPO, 2010; EPPO, 2014
Costa RicaPresentCABI/EPPO, 2010; EPPO, 2014
CubaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
CuraçaoPresentBen-Dov, 1993
DominicaPresentCABI/EPPO, 2010; EPPO, 2014
Dominican RepublicPresentCABI/EPPO, 2010; EPPO, 2014
GrenadaPresentCABI/EPPO, 2010; EPPO, 2014
GuadeloupePresentCABI/EPPO, 2010; EPPO, 2014
GuatemalaPresentCABI/EPPO, 2010; EPPO, 2014
HaitiPresentCABI/EPPO, 2010; EPPO, 2014
HondurasPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
JamaicaPresentCABI/EPPO, 2010; EPPO, 2014
MartiniquePresentCABI/EPPO, 2010; EPPO, 2014
MontserratPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
Netherlands AntillesPresentCABI/EPPO, 2010; EPPO, 2014
NicaraguaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
PanamaPresentCABI/EPPO, 2010; EPPO, 2014
Puerto RicoPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
Saint Kitts and NevisPresentCABI/EPPO, 2010; EPPO, 2014
Saint LuciaPresentCABI/EPPO, 2010; EPPO, 2014
Saint Vincent and the GrenadinesPresentCABI/EPPO, 2010; EPPO, 2014
Trinidad and TobagoPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
United States Virgin IslandsPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014

South America

BrazilPresentCABI/EPPO, 2010; EPPO, 2014
-Espirito SantoPresentCulik et al., 2007; CABI/EPPO, 2010; EPPO, 2014
-Minas GeraisPresentCABI/EPPO, 2010; EPPO, 2014
-Rio de JaneiroPresentCABI/EPPO, 2010; EPPO, 2014
-Rio Grande do SulPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-Sao PauloPresentCABI/EPPO, 2010; EPPO, 2014
ColombiaPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
EcuadorPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
GuyanaPresentCABI/EPPO, 2010; EPPO, 2014
VenezuelaPresentCABI/EPPO, 2010; EPPO, 2014

Europe

CyprusPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
FrancePresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-France (mainland)PresentCABI/EPPO, 2010
GreecePresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-CretePresentEPPO, 2014
ItalyPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
-Italy (mainland)PresentCABI/EPPO, 2010
MaltaPresentCABI/EPPO, 2010; EPPO, 2014
PortugalPresentCABI/EPPO, 2010; EPPO, 2014
-MadeiraPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
SpainAbsent, reported but not confirmedNHM, 1991; CABI/EPPO, 2010
-Spain (mainland)Absent, reported but not confirmedCABI/EPPO, 2010

Oceania

AustraliaPresentCABI/EPPO, 2010; EPPO, 2014
-New South WalesPresentCABI/EPPO, 2010; EPPO, 2014
-QueenslandPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
GuamPresentCABI/EPPO, 2010; EPPO, 2014
Micronesia, Federated states ofPresentCABI/EPPO, 2010; EPPO, 2014
Northern Mariana IslandsPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014
PalauPresentBen-Dov, 1993; CABI/EPPO, 2010; EPPO, 2014

Growth Stages

Top of page Flowering stage, Fruiting stage, Seedling stage, Vegetative growing stage

List of Symptoms/Signs

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SignLife StagesType
Fruit / honeydew or sooty mould
Growing point / external feeding
Leaves / abnormal colours
Leaves / abnormal leaf fall
Leaves / external feeding
Leaves / honeydew or sooty mould
Leaves / internal feeding
Leaves / wilting
Stems / external feeding
Stems / honeydew or sooty mould
Whole plant / external feeding

Natural enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Anicetus beneficus Parasite Adults/Nymphs Israel Citrus
Anicetus ceroplastis Parasite Adults/Nymphs
Aprostocetus ceroplastae Parasite Adults/Nymphs Egypt oranges
Blastobasis lecaniella Predator Adults/Nymphs
Cephaleta Parasite Adults/Nymphs
Cheiracanthium mildei Predator Adults/Nymphs Israel grapefruits
Cheletogenes ornatus Predator Adults/Nymphs
Chilocorus renipustulatus Predator Adults/Nymphs
Coccophagus ceroplastae Parasite Adults/Nymphs
Coccophagus hawaiiensis Parasite Adults/Nymphs Israel Citrus
Coccophagus lycimnia Parasite Adults/Nymphs
Cryptolaemus montrouzieri Predator
Cybocephalus micans Predator Adults/Nymphs
Diversinervus elegans Parasite
Eccopsia bodkini Predator Adults/Nymphs
Encarsia citrina Parasite Adults/Nymphs Egypt oranges
Metaphycus eruptor Parasite Adults/Nymphs
Metaphycus flavus Parasite Adults/Nymphs Egypt oranges
Metaseiulus occidentalis Predator Adults/Nymphs
Microterys clauseni Parasite Israel Citrus
Microterys flavus Parasite Egypt oranges
Microterys nietneri Parasite Adults/Nymphs
Microterys speciosus Parasite Israel Citrus
Moranila californica Predator Eggs
Scutellista caerulea Predator Eggs Israel Citrus
Vitula toboga Predator Adults/Nymphs

References

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Abbas MST, 1992. Comparative rates of infestation by four scale insects on citrus trees with special reference to rates of parasitism on the purple scale Lepidosaphes beckii. Egyptian Journal of Agricultural Research, 70(2):477-486; 7 ref.

Argov Y; Schneider B; Rosen D, 1992. Parasitism of Florida wax scale, Ceroplastes floridensis Comstock, on citrus in Israel. Journal of the Entomological Society of Southern Africa, 55(1):21-31

Argyriou LC; Kourmadas AL, 1980. Ceroplastes floridensis Comstock an important pest of citrus trees in pgean islands. Fruits, 35(11):705-708

Balachowsky AS, 1933. Sur la biologie de 'Ceroplastes floridensis' Comst. et sur la repartition géographique des Ceroplastes dans la region palearctique (Hem. Coccidae). Proceedings 5e Congres International d'Entomologie (Paris 1932), 5:79-87.

Ben-Dov Y, 1970. The wax scales of the genus Ceroplastes Gray (Homoptera: Coccidae) and their parasites in Israel. Israel Journal of Entomology, 5:83-92.

Ben-Dov Y, 1976. Phenology of the Florida wax scale, Ceroplastes floridensis Comstock (Homoptera: Coccidae), on citrus in Israel. Phytoparasitica, 4(1):3-7

Ben-Dov Y, 1993. A systematic catalogue of the soft scale insects of the world (Homoptera: Coccoidea: Coccidae) with data on geographical distribution, host plants, biology and economic importance. Gainesville, USA: Sandhill Crane Press, Inc., 536 pp.

Browning HW, 1994. Classical biological control of citrus scale insects. In Rosen D, Bennett FD, Capinera JL, eds. Pest management in the subtropics: biological control - a Florida perspective. Andover, UK: Intercept Limited, 49-78.

CABI/EPPO, 2010. Ceroplastes floridensis. [Distribution map]. Distribution Maps of Plant Pests, No.December. Wallingford, UK: CABI, Map 440 (1st revision).

CIE, 1982. Distribution Maps of Pests. No. 440. Wallingford, UK: CAB International.

Culik MP; Martins Ddos S; Ventura JA; Peronti ALBG; Gullan PJ; Kondo T, 2007. Coccidae, Pseudococcidae, Ortheziidae, and Monophlebidae(Hemiptera: Coccoidea) of Espírito Santo, Brazil. Biota Neotropica, 7(3):61-65. http://www.biotaneotropica.org.br/v7n3/en/fullpaper?bn00507032007+en

EPPO, 2014. PQR database. Paris, France: European and Mediterranean Plant Protection Organization. http://www.eppo.int/DATABASES/pqr/pqr.htm

Gimpel WF; Miller DR; Davidson JA, 1974. A systematic revision of the wax scales, genus Ceroplastes, in the United States (Homoptera; Coccoidea; Coccidae). Maryland, USA: University of Maryland, Agricultural Experiment Station, Miscellaneous Publication 841, 1-85.

Hamon AB; Williams ML, 1984. The soft scale insects of Florida (Homoptera: Coccoidea: Coccidae). Arthropods of Florida and Neighboring Land Areas, 11:1-94.

Helmy EI; Abdel-Megeed MI; El-Immry SM, 1991. Comparative potency of certain insecticides against Ceroplastes floridensis (Comst.). Egyptian Journal of Agricultural Research, 69(1):155-162; 15 ref.

Huang J; Huang BK, 1988. Bionomics of Ceroplastes floridensis Comstock (Hom. Coccidae) and its hymenopterous parasites (Hym., Chalcidoidea). Journal of Fujian Agricultural College, 17(1):31-37

Jendoubi H; Suma P; Russo A, 2012. First records of Ceroplastes floridensis Comstock (Hemiptera: Coccidae) and Chrysomphalus pinnulifer (Maskell) (Hemiptera: Diaspididae) in Tunisia. Entomological News, 122(3):247-249. http://www.bioone.org/loi/entn

Kawai S, 1980. Scale Insects of Japan in Colours. Tokyo, Japan: National Agriculture Education Association, 455 pp.

Kosztarab M, 1996. Scale Insects of North Eastern North America: identification, biology, and distribution. Martinsville, USA: Virginia Museum of Natural History, 650 pp.

Krombein KV, Hurd PD Jr. , Smith DR, Burks BD, 1979. Catalog of Hymenoptera in America North of Mexico. Washington DC, USA: Smithsonian Institution Press.

Mansour F; Whitecomb WH, 1986. The spiders of a citrus grove in Israel and their role as biocontrol agents of Ceroplastes floridensis (Homoptera: Coccidae). Entomophaga, 31(3):269-276

Peleg BA; Bar Zakay I; Ascher KRS; Ben-Dov Y, 1995. The pest status of citrus scale insects in Israel (1984-1994). Proceedings of the VII International Symposium of Scale Insect Studies, held in Bet Dagan, Israel, June 12-17 1994. Israel Journal of Entomology, 29:261-264.

Pellizzari G; Camporese P, 1994. The Ceroplastes species (Homoptera: Coccoidea) of the Mediterranean Basin with emphasis on C. japonicus Green. Annales de la Societe Entomologique de France, 30(2):175-192

Podoler H; Dreishpoun Y; Rosen D, 1981. Population dynamics of the Florida wax scale, Ceroplastes floridensis (Homoptera: Coccidae) on citrus in Israel. 1. A partial life-table. Acta Oecologica, Oecologia Applicata, 2(1):81-91

Salem SA; Zaki FN, 1984. Economic threshold level of the citrus wax scale Ceroplastes floridensis Comstock on citrus, Egypt. Bulletin de la Societe Entomologique d'Egypte, No. 65:333-344

Schneider B; Podoler H; Rosen D, 1987. Population dynamics of the Florida wax scale, Ceroplastes floridensis (Homoptera: Coccidae), on citrus in Israel. 2. Spatial distribution. Acta Oecologica, Oecologia Applicata, 8(1):67-78

Sun JZ; Qian ZG; Liu FL; Song HW; Wen TH; Ma PY; Wang YC; Li YL, 1989. Study on the techniques for integrated control of the major diseases and insects of Zizyphus jujuba. Forest Science and Technology, No. 4:1-4

Swailem SM; Ismail II; Ahmed NMAS, 1976. Population studies on the citrus wax scale insect, Ceroplastes floridensis Comstock, in different regions in Egypt (Homoptera: Coccidae). Bulletin of the Entomological Society of Egypt, No. 60:229-237

Tang FT, 1991. The Coccidae of China. Taiyuan, China: Shanxi United Universities Press.

Vidyasagar PSPV; Koya KMA; Devasahayam S; Premkumar T, 1989. Record of wax scale Ceroplastes floridensis Comstock (Homoptera: Coccidae) infesting clove seedlings in Kerala, India. Entomon, 14(3-4):359-360

Williams DJ; Watson GW, 1990. The scale insects of the tropical South Pacific region. Part 3: the soft scales (Coccidae) and other families. Wallingford, UK: CAB International, 267 pp.

Williams ML; Kosztarab M, 1972. Morphology and systematics of the Coccidae of Virginia with notes on their biology (Homoptera: Coccoidea). Research Division Bulletin of the Virginia Polytechnic Institute and State University, Blacksburg, 74:1-215.

Yardeni A, 1987. Evaluation of wind dispersed soft scale crawlers (Homoptera: Coccidae), in the infestation of a citrus grove in Israel. Israel Journal of Entomology, 21:25-31.

Yardeni A; Rosen D, 1990. Wind dispersal and pattern of colonization of Ceroplastes floridensis on citrus in Israel. In: Proceedings of the Sixth International Symposium of Scale Insect Studies, Part II. Cracow, Poland, August 6-12, 1990. Cracow, Poland: Agricultural University Press, 125-128.

Yardeni A; Shapira E; Ascher KRS; Ben-Dov Y, 1995. Thinning populations of the Florida wax scale, Ceroplastes floridensis Comstock (Coccidae), by use of potassium nitrate and spray oil, as an option in IPM of citrus groves in Israel. Proceedings of the VII International Symposium of Scale Insect Studies, held in Bet Dagan, Israel, June 12-17 1994. Israel Journal of Entomology, 29:271-276.

Ülgentürk S; Sahin Ö; Ayhan B; Saribasak H; Kaydan MB, 2012. Coccoidea (Hemiptera) species of Taurus cedar (Cedrus libani) in Turkey. (Türkiye'de Toros sedirinin (Cedrus libani) Coccoidea (Hemiptera) türleri.) Türkiye Entomoloji Dergisi, 36(1):113-121. http://entomoloji.ege.edu.tr/files/Arsiv/2012_36_1/2012_36_1_113-121.pdf

Distribution Maps

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