Cyanthillium cinereum (little ironweed)
Index
- Pictures
- Identity
- Summary of Invasiveness
- Taxonomic Tree
- Notes on Taxonomy and Nomenclature
- Description
- Plant Type
- Distribution
- Distribution Table
- History of Introduction and Spread
- Risk of Introduction
- Habitat
- Habitat List
- Hosts/Species Affected
- Host Plants and Other Plants Affected
- Growth Stages
- Biology and Ecology
- Climate
- Rainfall Regime
- Soil Tolerances
- Means of Movement and Dispersal
- Pathway Causes
- Pathway Vectors
- Impact Summary
- Economic Impact
- Threatened Species
- Risk and Impact Factors
- Uses
- Uses List
- Similarities to Other Species/Conditions
- References
- Links to Websites
- Contributors
- Distribution Maps
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Top of pagePreferred Scientific Name
- Cyanthillium cinereum (L.) H. Rob
Preferred Common Name
- little ironweed
Other Scientific Names
- Blumea esquirolii H.Lev. & Vaniot
- Cacalia arguta Kuntze
- Cacalia cinerea (L.) Kuntze
- Cacalia erigerodes Kuntze
- Cacalia exilis Kuntze
- Cacalia kroneana Kuntze
- Cacalia linfolia DC.
- Cacalia rotundifolia Willd.
- Cacalia vialis Kuntze
- Calea cordata Lour.
- Cineraria glaberrima Spreng. ex DC.
- Conyza chinensis L.
- Conyza cinerea L.
- Conzya heterophylla Lam.
- Conzya incana DC.
- Conzya prolifera Lam.
- Crassocephalum flatmense Hochst. & Steud. ex DC.
- Cyanopis decurrens Zoll. & Mor.
- Eupatorium arboreum Reinw. ex de Vriese
- Eupatorium myosotifolium Jacq.
- Pteronia tomentosa Lour.
- Seneciodes cinereum (L.) Kuntze
- Serratula cinerea (L.) Roxb.
- Vernonia abbreviata DC.
- Vernonia arguta Baker
- Vernonia betonicaefolia Baker
- Vernonia cinerea (L.) Less.
- Vernonia exilis Miq.
- Vernonia fasciculata Blume
- Vernonia kroneana Miq.
- Vernonia vialis DC.
International Common Names
- English: ironweed; small ironweed; vernonia
- Spanish: machadita; rabo de buey; yerba morada
- French: ayapana sauvage
- Chinese: ye xiang niu
Local Common Names
- Cuba: machadita
- Fiji: kaukamea; tho vuka; vutikaumondro
- India: ankari; ankta; pokasubgo; sahaderi; sandri
- Indonesia: maryuna
- Japan: mura-saki-mukashi-yomogi; reinan-nogiku; yambaru-higotai
- Lesser Antilles: inflammation bush; iron bush; measle bush
- Malaysia: rumput taki babi; tambak-tambak
- Mauritius: ayapa sauvage
- Nigeria: bojure
- Philippines: agas-moro; bulak-manok; kolong-kugon
- Puerto Rico: rabo de buey; yerba socialista
- Sri Lanka: alavangu pillu; monara kudumbiya
- Thailand: kaan thuup; yaa dok khaao; yaa-saam-wan; ya-la-ong
Summary of Invasiveness
Top of pageC. cinereum is a cosmopolitan weed common in disturbed areas in tropical and subtropical regions of the world (Randall, 2012). It is a fast-growing, annual herb with the capacity to form dense patches in gardens, roadsides, waste grounds and pasture (Holm et al., 1997). Currently, this species is considered invasive in many islands in the Pacific Ocean (e.g. Hawaii, Fiji, French Polynesia and Micronesia), New Zealand, Singapore, Costa Rica, Guatemala, Nicaragua, Panama, Galápagos Islands, Cuba, Puerto Rico and the Virgin Islands (see distribution table for details; Chong et al., 2009; Chacon and Saborio, 2012; González-Torres et al., 2012; PIER, 2013).
Taxonomic Tree
Top of page- Domain: Eukaryota
- Kingdom: Plantae
- Phylum: Spermatophyta
- Subphylum: Angiospermae
- Class: Dicotyledonae
- Order: Asterales
- Family: Asteraceae
- Genus: Cyanthillium
- Species: Cyanthillium cinereum
Notes on Taxonomy and Nomenclature
Top of pageThe family Asteraceae is one of the most diverse groups among flowering plants including 1620 genera and about 23,600 species (Stevens, 2012). Species in the Asteraceae are very variable vegetatively, but may be recognized by their “capitulate” and involucrate inflorescences in which numerous small flowers open first on the outside and are infrequently subtended by bracts. The anthers in this family are usually fused and form a tube through which the style extends before the two stigmatic lobes separate and become recurved. The rather small, single-seeded fruits usually have a plumose “pappus” and are frequently dispersed by wind (Stevens, 2012).
The genus Cyanthillium has been used as a synonym of Vernonia, while other botanists have treated both genera as distinct. Currently, both genera are treated separately and Cyanthillium is distinguished from Vernonia by its smooth to finely 5-striate (vs. 5-10-ribbed) achenes, and by its echinolophate (vs. tricolporate) pollen, and by base chromosome number usually reported as 2n = 18 or 20 (vs. 2n = 34). Currently five species are recognized within Cyanthillium (Funk and Pruski, 1996).
Description
Top of pageAnnual herb, to 80 (-150) cm tall. Stems unbranched or more commonly few-branched. Leaves petiolate or distal ones sessile; petioles to 1.5 cm, narrowly winged; blade 1.5-5 × 1-1.5(4) cm, obovate or less often elliptic or ovate, base cuneate, apex acute to obtuse, lower surface pilose, usually glandular. Inflorescence of 5 to numerous heads, 13-20-flowered; involucre 2.5-3 mm long; outer phyllaries much reduced, inner phyllaries subequal. Corolla 3-4 mm, exserted 1.5-2 mm from involucre, the tube long and narrow, 2-3 mm, the limbs short, pilose. Achenes 1.2–2 mm long, subfusiform, terete not ribbed, inner pappus white, exserted from involucre and nearly as long as the corollas (Funk and Pruski, 1996).
Distribution
Top of pageC. cinereum is native to Africa, tropical and temperate Asia, and Australia (USDA-ARS, 2013). It is widely naturalized in tropical regions in America and islands in the Pacific (PIER, 2013; USDA-ARS, 2013).
Distribution Table
Top of pageThe distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.
Last updated: 10 Jan 2020Continent/Country/Region | Distribution | Last Reported | Origin | First Reported | Invasive | Reference | Notes |
---|---|---|---|---|---|---|---|
Africa |
|||||||
Benin | Present | Native | |||||
Burundi | Present | Native | |||||
Cameroon | Present | Native | |||||
Central African Republic | Present | Native | |||||
Congo, Democratic Republic of the | Present | Native | |||||
Côte d'Ivoire | Present | Native | |||||
Eritrea | Present | Native | |||||
Eswatini | Present | Native | |||||
Ethiopia | Present | Native | |||||
Gabon | Present | Native | |||||
Ghana | Present | Native | |||||
Guinea | Present | Native | |||||
Guinea-Bissau | Present | Native | |||||
Kenya | Present | Native | |||||
Liberia | Present | Native | |||||
Madagascar | Present | Native | |||||
Mali | Present | Native | |||||
Mayotte | Present | Introduced | Invasive | ||||
Niger | Present | Native | |||||
Réunion | Present | Native | |||||
Rwanda | Present | Native | |||||
Senegal | Present | Native | |||||
Seychelles | Present | Introduced | Invasive | ||||
Sierra Leone | Present | Native | |||||
Somalia | Present | Native | |||||
South Africa | Present | Native | |||||
Tanzania | Present | Native | |||||
Togo | Present | Native | |||||
Uganda | Present | Native | |||||
Asia |
|||||||
Bangladesh | Present | Weed | |||||
Bhutan | Present | Native | |||||
British Indian Ocean Territory | |||||||
-Chagos Archipelago | Present | Introduced | Invasive | ||||
China | Present | Present based on regional distribution. | |||||
-Fujian | Present | Native | Considered a weed | ||||
-Guangdong | Present | Native | Considered a weed | ||||
-Guangxi | Present | Native | Considered a weed | ||||
-Hubei | Present | Native | Considered a weed | ||||
-Hunan | Present | Native | Considered a weed | ||||
-Jiangxi | Present | Native | Considered a weed | ||||
-Sichuan | Present | Native | Considered a weed | ||||
-Yunnan | Present | Native | Considered a weed | ||||
-Zhejiang | Present | Native | Considered a weed | ||||
India | Present | Native | |||||
Indonesia | Present | Native | |||||
Japan | Present | Native | |||||
-Kyushu | Present | Native | |||||
-Ryukyu Islands | Present | Native | |||||
Malaysia | Present | Native | |||||
Maldives | Present | ||||||
Myanmar | Present | Native | |||||
Oman | Present | Native | |||||
Philippines | Present | Native | |||||
Singapore | Present | Invasive | |||||
Sri Lanka | Present | Native | |||||
Taiwan | Present | Native | Considered a weed | ||||
Thailand | Present | Native | |||||
Vietnam | Present | Native | |||||
Yemen | Present | Native | |||||
North America |
|||||||
Anguilla | Present | Introduced | |||||
Antigua and Barbuda | Present | Introduced | |||||
Bahamas | Present | Introduced | |||||
Barbados | Present | Introduced | |||||
Belize | Present | Introduced | |||||
British Virgin Islands | Present | Introduced | Guana, Tortola, Virgin Gorda | ||||
Costa Rica | Present | Introduced | Invasive | ||||
Cuba | Present | Introduced | Invasive | Highly invasive | |||
Dominica | Present | Introduced | |||||
Dominican Republic | Present | Introduced | |||||
Grenada | Present | Introduced | |||||
Guatemala | Present | Introduced | Invasive | ||||
Haiti | Present | Introduced | |||||
Honduras | Present | Introduced | Invasive | ||||
Jamaica | Present | Introduced | |||||
Martinique | Present | Introduced | |||||
Mexico | Present | Introduced | Weed | ||||
Montserrat | Present | Introduced | |||||
Netherlands Antilles | Present | Introduced | Saba, St. Marteen | ||||
Nicaragua | Present | Introduced | Invasive | ||||
Panama | Present | Introduced | Invasive | ||||
Puerto Rico | Present | Introduced | Invasive | ||||
Saint Kitts and Nevis | Present | Introduced | |||||
Saint Lucia | Present | Introduced | |||||
Saint Vincent and the Grenadines | Present | Introduced | |||||
Trinidad and Tobago | Present | Introduced | |||||
U.S. Virgin Islands | Present | Introduced | Invasive | ||||
United States | Present | Present based on regional distribution. | |||||
-Florida | Present | Introduced | |||||
-Hawaii | Present | Introduced | Invasive | ||||
Oceania |
|||||||
American Samoa | Present | Introduced | Invasive | ||||
Australia | Present | Present based on regional distribution. | |||||
-New South Wales | Present | Native | |||||
-Northern Territory | Present | Native | |||||
-Queensland | Present | Native | |||||
-Western Australia | Present | Native | |||||
Christmas Island | Present | Introduced | Invasive | ||||
Cook Islands | Present | Introduced | |||||
Federated States of Micronesia | Present | Invasive | This species has been listed as native for this archipelago | ||||
Fiji | Present | Introduced | Invasive | ||||
French Polynesia | Present | Introduced | Invasive | ||||
Guam | Present | Introduced | Invasive | ||||
Kiribati | Present | Introduced | Invasive | Formerly listed as native | |||
Marshall Islands | Present | Invasive | Listed as native for some islands and naturalised and invasive for other islands | ||||
Nauru | Present | Introduced | Invasive | Formerly listed as native | |||
New Caledonia | Present | Introduced | Invasive | ||||
Niue | Present | Introduced | Invasive | ||||
Northern Mariana Islands | Present | Invasive | Listed as native for some islands and naturalised and invasive for other islands | ||||
Palau | Present | Native | |||||
Papua New Guinea | Present | Native | |||||
Pitcairn | Present | Introduced | Invasive | ||||
Samoa | Present | ||||||
Solomon Islands | Present | ||||||
Tonga | Present | Introduced | Invasive | ||||
Tuvalu | Present | ||||||
U.S. Minor Outlying Islands | Present | Introduced | Invasive | ||||
Vanuatu | Present | ||||||
Wallis and Futuna | Present | Introduced | Invasive | ||||
South America |
|||||||
Brazil | Present | Present based on regional distribution. | |||||
-Amazonas | Present | Introduced | |||||
-Bahia | Present | Introduced | |||||
-Minas Gerais | Present | Introduced | |||||
-Para | Present | Introduced | |||||
-Pernambuco | Present | Introduced | |||||
-Rio de Janeiro | Present | Introduced | |||||
-Sao Paulo | Present | Introduced | |||||
Ecuador | Present | Present based on regional distribution. | |||||
-Galapagos Islands | Present | Introduced | Invasive | ||||
French Guiana | Present | Introduced | Naturalized | Naturalised | |||
Guyana | Present | Introduced | Naturalized | Naturalised | |||
Suriname | Present | Introduced | Naturalized | Naturalised | |||
Venezuela | Present | Introduced | Naturalized | Naturalised |
History of Introduction and Spread
Top of pageThe history of introduction of C. cinereum is uncertain. It is highly probable that the species has been introduced accidentally as a contaminant or as a weed in nursery materials (Holm et al., 1997). For the West Indies, herbarium collections show that it was first collected in Jamaica in 1882 (US Herbarium Collection). In Central America the date of introduction is unclear but since the 1950s the species appears listed as a weed (Cardenas and Coulston, 1967).
Risk of Introduction
Top of pageThe risk of introduction of C. cinereum is high. This species is easily dispersed by wind and has the potential to grow as a weed in ruderal areas, and agricultural and pasture lands. C. cinereum has been reported as a weed in 27 crops in 47 countries in Asia, East and West Africa and the Caribbean (Holm et al., 1997). Consequently, this species has the potential to spread much further than it has to date.
Habitat
Top of pageC. cinereum is a common weed in coastal areas, beaches, cultivated areas, disturbed areas, pastures, roadsides, rocky areas, savannas, secondary vegetation, and gardens. In Africa (within its native distribution range), this species grows at lower altitudes (0-800 m) in woodlands and grasslands (PROTA4U, 2013).
Habitat List
Top of pageCategory | Sub-Category | Habitat | Presence | Status |
---|---|---|---|---|
Terrestrial | ||||
Terrestrial | Managed | Cultivated / agricultural land | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Managed | Cultivated / agricultural land | Present, no further details | Natural |
Terrestrial | Managed | Managed forests, plantations and orchards | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Managed | Managed forests, plantations and orchards | Present, no further details | Natural |
Terrestrial | Managed | Managed grasslands (grazing systems) | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Managed | Managed grasslands (grazing systems) | Present, no further details | Natural |
Terrestrial | Managed | Disturbed areas | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Managed | Disturbed areas | Present, no further details | Natural |
Terrestrial | Managed | Rail / roadsides | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Managed | Rail / roadsides | Present, no further details | Natural |
Terrestrial | Managed | Urban / peri-urban areas | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Managed | Urban / peri-urban areas | Present, no further details | Natural |
Terrestrial | Natural / Semi-natural | Natural forests | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Natural / Semi-natural | Natural forests | Present, no further details | Natural |
Terrestrial | Natural / Semi-natural | Natural grasslands | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Natural / Semi-natural | Natural grasslands | Present, no further details | Natural |
Littoral | Coastal areas | Present, no further details | Harmful (pest or invasive) | |
Littoral | Coastal areas | Present, no further details | Natural |
Hosts/Species Affected
Top of pageC. cinereum is reported as a weed in 27 crops in 47 countries (Holm et al., 1997).
Serious weed in:
- sugarcane, cotton, groundnuts and wheat in India
- pastures in Australia, India, Nigeria and Thailand
- rice in Philippines;
- taro in Samoa.
Common weed in:
- banana in Surinam and Tonga
- cassava in Surinam
- cocoa in Indonesia
- citrus in Surinam
- cotton in the Philippines
- maize in India
- oil palm in Surinam
- pastures in Australia, Dominican Republic, Puerto Rico and Jamaica
- pineapple in Hawaii
- rice in Surinam, India, Indonesia and Sri Lanka
- rubber in Indonesia and Thailand
- sugarcane in Bangladesh, Hawaii, and the Philippines
- taro in Tonga
- tea in India and Indonesia
- vegetables in Surinam and Thailand
Unranked weed in:
- abaca (Musa textilis) in the Philippines
- cocoa in Dominican Republic
- cassava in India, Indonesia, and Nigeria
- coconut in Sri Lanka and Surinam
- coffee in Dominican Republic
- cotton in Mozambique and Tanzania
- legumes and tomatoes in the Philippines
- macadamia nut in Hawaii
- maize in Cambodia, Gambia, Indonesia, Nigeria, the Philippines and Zambia
- pastures in the Philippines
- groundnuts in Indonesia and Nigeria
- rice in Laos, Thailand and Vietnam
- rubber and tea in Sri Lanka
- sugarcane in British Guiana, Dominican Republic, Laos and Vietnam
- tobacco in the Philippines
Host Plants and Other Plants Affected
Top of pagePlant name | Family | Context | References |
---|---|---|---|
Ananas comosus (pineapple) | Bromeliaceae | Main | |
Arachis hypogaea (groundnut) | Fabaceae | Main | |
Citrus | Rutaceae | Main | |
Cocos nucifera (coconut) | Arecaceae | Main | |
Coffea arabica (arabica coffee) | Rubiaceae | Main | |
Colocasia esculenta (taro) | Araceae | Main | |
Elaeis guineensis (African oil palm) | Arecaceae | Main | |
Gossypium (cotton) | Malvaceae | Main | |
Hevea brasiliensis (rubber) | Euphorbiaceae | Main | |
Macadamia integrifolia (macadamia nut) | Proteaceae | Main | |
Manihot esculenta (cassava) | Euphorbiaceae | Main | |
Musa (banana) | Musaceae | Main | |
Nicotiana | Solanaceae | Main | |
Oryza sativa (rice) | Poaceae | Main | |
Saccharum | Poaceae | Main | |
Solanum lycopersicum (tomato) | Solanaceae | Main | |
Theobroma cacao (cocoa) | Malvaceae | Main | |
Triticum (wheat) | Poaceae | Main | |
Zea mays (maize) | Poaceae | Main |
Biology and Ecology
Top of pageGenetics
C. cinereum plants are diploid with a chromosome number of 2n = 18 (Holm et al., 1997; Pruski 2013).
Reproductive Biology
Flowers in C. cinereum are pollinated by wind. Probably the most common reproduction system in this genus is allogamy with a sporophytic self-incompatibility (Holm et al., 1997; Flora of China Editorial Committee, 2012).
Physiology and Phenology
C. cinereum is an annual herb and under favourable environmental conditions it produces flowers and seeds for many months (Holm et al., 1997; Flora of China Editorial Committee, 2012).
Environmental Requirements
C. cinereum usually grows as a weed, thus it needs full sunlight and moderate water availability to grow. It prefers sandy-loam soils but can be found growing on a range of soils with pH ranging from 4 to 6. It is able to tolerate semiarid conditions as well as partial salinity conditions (PROTA4U, 2013).
Climate
Top of pageClimate | Status | Description | Remark |
---|---|---|---|
Af - Tropical rainforest climate | Preferred | > 60mm precipitation per month | |
Am - Tropical monsoon climate | Preferred | Tropical monsoon climate ( < 60mm precipitation driest month but > (100 - [total annual precipitation(mm}/25])) | |
As - Tropical savanna climate with dry summer | Preferred | < 60mm precipitation driest month (in summer) and < (100 - [total annual precipitation{mm}/25]) | |
Aw - Tropical wet and dry savanna climate | Preferred | < 60mm precipitation driest month (in winter) and < (100 - [total annual precipitation{mm}/25]) | |
BS - Steppe climate | Tolerated | > 430mm and < 860mm annual precipitation | |
Cs - Warm temperate climate with dry summer | Tolerated | Warm average temp. > 10°C, Cold average temp. > 0°C, dry summers | |
Cw - Warm temperate climate with dry winter | Tolerated | Warm temperate climate with dry winter (Warm average temp. > 10°C, Cold average temp. > 0°C, dry winters) |
Soil Tolerances
Top of pageSoil drainage
- free
Soil reaction
- acid
- neutral
Soil texture
- heavy
- light
- medium
Special soil tolerances
- saline
Means of Movement and Dispersal
Top of pageC. cinereum spreads by seeds which are adapted to wind dispersal. In addition, seeds may be secondarily dispersed as a contaminant in crop seeds, pasture seeds and in agricultural machinery (Holm et al., 1997; Pruski 2013).
Pathway Causes
Top of pageCause | Notes | Long Distance | Local | References |
---|---|---|---|---|
Crop production | Probably dispersed as a seed and crop contaminant | Yes | Yes | Holm et al., 1997 |
Disturbance | Common weed | Yes | Yes | Holm et al., 1997 |
Escape from confinement or garden escape | Common weed | Yes | Yes | Holm et al., 1997 |
Forage | Common weed in pasturelands | Yes | Yes | Holm et al., 1997 |
Garden waste disposal | Common weed in gardens | Yes | Yes | Holm et al., 1997 |
Horticulture | Common weed | Yes | Yes | Holm et al., 1997 |
Medicinal use | Used in African traditional herb medicine | Yes | Yes | Prota4U, 2013 |
Pathway Vectors
Top of pageVector | Notes | Long Distance | Local | References |
---|---|---|---|---|
Machinery and equipment | Seeds | Yes | Yes | Holm et al., 1997 |
Soil, sand and gravel | Seeds | Yes | Yes | Holm et al., 1997 |
Water | Seeds | Yes | Yes | Holm et al., 1997 |
Impact Summary
Top of pageCategory | Impact |
---|---|
Economic/livelihood | Negative |
Environment (generally) | Negative |
Human health | Positive and negative |
Economic Impact
Top of pageC. cinereum is a weed with important economic impacts mainly in agriculture, pasture and garden activities. It is listed as a weed in about 27 different crops in 47 countries in Asia, Australia, Africa and America. In addition, C. cinereum has the potential to negatively impact natural and seminatural habitats in coastal areas, forest edges, secondary forests, upland forests, and grasslands (Pruski, 2013).
C. cinereum can host the tobacco leaf curl virus and the root-knot nematode (Holm et al., 1997).
Threatened Species
Top of pageThreatened Species | Conservation Status | Where Threatened | Mechanism | References | Notes |
---|---|---|---|---|---|
Schiedea spergulina var. leiopoda | National list(s); USA ESA listing as endangered species | Hawaii | Competition - monopolizing resources | US Fish and Wildlife Service, 2010 |
Risk and Impact Factors
Top of page- Invasive in its native range
- Proved invasive outside its native range
- Has a broad native range
- Abundant in its native range
- Highly adaptable to different environments
- Is a habitat generalist
- Tolerates, or benefits from, cultivation, browsing pressure, mutilation, fire etc
- Pioneering in disturbed areas
- Highly mobile locally
- Benefits from human association (i.e. it is a human commensal)
- Fast growing
- Damaged ecosystem services
- Ecosystem change/ habitat alteration
- Loss of medicinal resources
- Modification of successional patterns
- Monoculture formation
- Negatively impacts agriculture
- Negatively impacts cultural/traditional practices
- Negatively impacts forestry
- Competition - monopolizing resources
- Pest and disease transmission
- Rapid growth
- Highly likely to be transported internationally accidentally
- Difficult to identify/detect as a commodity contaminant
- Difficult/costly to control
Uses
Top of pageLeaves of C. cinereum have some antibiotic activity and consequently they are frequently used in traditional medicine in India to treat conjunctivitis, dropsy and urinary disorders. In the Philippines, this species is used as an infusion for cough and skin diseases. A poultice from leaves reduces headaches while a root decoction relieves stomach aches and diarrhea. In Thailand, the leaves are used in the treatment of asthma and bronchitis (Holm et al., 1997; PROTA4U, 2013). The young shoots are eaten as a cooked vegetable in Java (PROTA4U, 2013).
Similarities to Other Species/Conditions
Top of pageSpecies of Cyanthillium can be confused with species in the genus Vernonia. These two genera can be distinguished by the following characters:
- In Cyanthillium the cypselae is smooth to finely 5-striate while in Vernonia cypselae is 5-10-ribbed.
- Pollen in Cyanthillium is echinolophate while in Vernonia pollen is tricolporate
- The chromosome number usually reported for Cyanthillium species is 2n = 18 or 20 and for Vernonia species is 2n = 34.
The species C. cinereum is recognized by the slender, ribbed stem, shallowly toothed leaves and the branched corymbs with bright purple to pinkish violet flower heads (Holm et al., 1997).
References
Top of pageAcevedo-Rodríguez P, Strong MT, 2012. Catalogue of the Seed Plants of the West Indies. Smithsonian Contributions to Botany, 98:1192 pp. Washington DC, USA: Smithsonian Institution. http://botany.si.edu/Antilles/WestIndies/catalog.htm
Balick MJ, Nee M, Atha DE, 2000. Checklist of the vascular plants of Belize. Memoirs of the New York Botanical Garden, 85:1-246.
Cardenas J, Coulston L, 1967. Weeds: A List of Common and Scientific Names for Brazil, Colombia, Puerto Rico, and Central America.
Chacón E, Saborío G, 2012. Red Interamericana de Información de Especies Invasoras, Costa Rica ([English title not available]). San José, Costa Rica: Asociación para la Conservación y el Estudio de la Biodiversidad. http://invasoras.acebio.org
Charles Darwin Foundation, 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Galapagos, Ecuador: Charles Darwin Foundation.
Chong KY, Tan HTW, Corlett RT, 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species., Singapore: Raffles Museum of Biodiversity Research, National University of Singapore, 273 pp.
Flora of China Editorial Committee, 2012. Flora of China Web. Cambridge, USA: Harvard University Herbaria. http://flora.huh.harvard.edu/china/
Florence J, Chevillotte H, Ollier C, Meyer J-Y, 2013. Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP) (Botanical database of the Nadeaud Herbarium of French Polynesia). http://www.herbier-tahiti.pf
Forzza RC, Leitman PM, Costa AF, Carvalho Jr AA, et al. , 2012. List of species of the Flora of Brazil (Lista de espécies Flora do Brasil). Rio de Janeiro, Brazil: Rio de Janeiro Botanic Garden. http://floradobrasil.jbrj.gov.br/2012/
Fosberg FR, Stoddart DR, 1994. Flora of the Phoenix Islands, central Pacific. Atoll Research Bulletin, 393:60 pp.
Funk VA, Pruski J, 1996. Asteraceae. Memoirs of the New York Botanical Garden, 78:85-122. [Flora of St. John, US Virgin Islands.]
González-Torres LR, Rankin R, Palmarola A (eds), 2012. Invasive plants in Cuba. (Plantas Invasoras en Cuba.) Bissea: Boletin sobre Conservacion de Plantad del Jardin Botanico Nacional, 6:1-140.
Graveson R, 2012. Plants of Saint Lucia. http://www.saintlucianplants.com
Herrera K, Lorence D, Flynn T, Balick MJ, 2010. Checklist of the vascular plants of Pohnpei with local names and uses. Lawai, Hawaii, USA: National Tropical Botanical Garden, 146 pp.
Hokche O, Berry PE, Huber O, 2008. Nuevo Catálogo de la Flora Vascular de Venezuela (New catalogue of the vascular flora of Venezuela). Caracas, Venezuela: Fundación Instituto Botánico de Venezuela, 860 pp.
Holm L, Doll J, Holm E, Pancho J, Herberger J, 1997. World Weeds. Natural Histories and Distribution. New York, USA: John Wiley and Sons, Inc.
Madagascar Catalogue, 2012. Catalogue of the vascular plants of Madagascar. Missouri Botanical Garden. St Louis, USA: Missouri Botanical Garden and Madagascar Research and Conservation Program, Antananarivo. http://www.efloras.org/madagascar
McCormack G, 2013. Cook Islands Biodiversity Database, Version 2007. Cook Islands Biodiversity Database. Rarotonga, Cook Islands: Cook Islands Natural Heritage Trust. http://cookislands.bishopmuseum.org/search.asp
Meyer JY, 2007. Rapport de mission sur l'Ile d'Uvea (Wallis & Futuna) du 6 au 17 Novembre 2007: Inventaire preliminaire de la flore vasculaire secondaire ([English title not available]). Papeete, Tahiti: Ministère de l'Education, l'Enseignement Supérieur et la Recherche, 39 pp. http://www.li-an.fr/jyves/Meyer_2007_Rapport_Plantes_Introduites_Wallis.pdf
Orchard AE, 1993. Flora of Australia. Vol. 50, Oceanic islands 2. Canberra: Australian Government Publishing Service.
PIER, 2013. Pacific Islands Ecosystems at Risk. Honolulu, Hawaii, USA: HEAR, University of Hawaii. http://www.hear.org/pier/index.html
Prota4U, 2013. PROTA4U web database. Grubben GJH, Denton OA, eds. Wageningen, Netherlands: Plant Resources of Tropical Africa. http://www.prota4u.org/search.asp
Pruski JF, 2013. Asteraceae. In: Flora Mesoamericana, Volume 5 (2) [ed. by Davidse, G. \Sousa Sánchez, M. \Knapp, S. \Chiang Cabrera, F.]., Mexico: Universidad Nacional Autónoma de México. http://www.tropicos.org/docs/meso/asteraceae.pdf
Randall RP, 2012. A Global Compendium of Weeds. Perth, Australia: Department of Agriculture and Food Western Australia, 1124 pp. http://www.cabi.org/isc/FullTextPDF/2013/20133109119.pdf
Smith AC, 1991. Flora Vitiensis nova: A new flora of Fiji. Lawai, Kauai, Hawai`i. National Tropical Botanical Garden, Volume 5, 626 pp.
Space JC, Waterhouse BM, Newfield M, Bull C, 2004. Report to the Government of Niue and the United Nations Development Programme: Invasive Plant Species on Niue following Cyclone Heta. 76 pp. http://www.hear.org/pier/pdf/niue_report_20041217.pdf
Stevens PF, 2012. Angiosperm Phylogeny Website. http://www.mobot.org/MOBOT/research/APweb/
Stone BC, 1970. The Flora of Guam. Micronesica, 6:1-659.
Thaman RR, Fosberg FR, Manner HI, Hassall DC, 1994. The flora of Nauru. Atoll Research Bulletin, 392:1-223.
USDA-ARS, 2013. Germplasm Resources Information Network (GRIN). Online Database. Beltsville, Maryland, USA: National Germplasm Resources Laboratory. https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysearch.aspx
Vibrans H, 2009. Malezas de México- Pennisetum purpureum ([English title not available]). http://www.conabio.gob.mx/malezasdemexico/poaceae/pennisetum-purpureum/fichas/ficha.htm
Wagner WL, Herbst DR, Sohmer SH, 1999. Manual of the flowering plants of Hawaii. Revised edition. Honolulu, Hawaii, USA: University of Hawaii Press/Bishop Museum Press. [Bernice P. Bishop Museum special publication.]
Wagner WL, Herbst DR, Weitzman A, Lorence DH, 2013. Flora of Micronesia. Flora of Micronesia (online). National Tropical Botanical Garden and the Smithsonian Institution. http://botany.si.edu/pacificislandbiodiversity/micronesia/index.htm
Whistler WA, 1983. The flora and vegetation of Swains Island. Atoll Research Bulletin, 262:25 pp.
Whistler WA, 1996. Botanical survey of Diego Garcia, Chagos Archipelago, British Indian Ocean Territory. Isle Botanica (online), 49 pp. http://www.zianet.com/tedmorris/dg/2005NRMP-Appendixe-botanicalsurvey.pdf
Distribution References
Anon, 2012. Invasive plants in Cuba. (Plantas Invasoras en Cuba). In: Bissea: Boletin sobre Conservacion de Plantad del Jardin Botanico Nacional, 6 [ed. by González-Torres LR, Rankin R, Palmarola A]. 1-140.
Balick MJ, Nee M, Atha DE, 2000. Checklist of the vascular plants of Belize. In: Memoirs of the New York Botanical Garden, 85 1-246.
CABI, Undated. CABI Compendium: Status inferred from regional distribution. Wallingford, UK: CABI
CABI, Undated a. CABI Compendium: Status as determined by CABI editor. Wallingford, UK: CABI
Chacón E, Saborío G, 2012. [English title not available]. (Red Interamericana de Información de Especies Invasoras, Costa Rica)., San José, Costa Rica: Asociación para la Conservación y el Estudio de la Biodiversidad. http://invasoras.acebio.org
Charles Darwin Foundation, 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos., Galapagos, Ecuador: Charles Darwin Foundation.
Chong KY, Tan HTW, Corlett RT, 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species., Singapore: Raffles Museum of Biodiversity Research, National University of Singapore. 273 pp.
Flora of China Editorial Committee, 2012. Flora of China Web., Cambridge, USA: Harvard University Herbaria. http://flora.huh.harvard.edu/china/
Florence J, Chevillotte H, Ollier C, Meyer JY, 2013. Botanical database of the Nadeaud Herbarium of French Polynesia. (Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP))., http://www.herbier-tahiti.pf
Forzza RC, Leitman PM, Costa AF, Carvalho Jr AA et al, 2012. List of species of the Flora of Brazil. (Lista de espécies Flora do Brasil)., Rio de Janeiro, Brazil: Rio de Janeiro Botanic Garden. http://floradobrasil.jbrj.gov.br/2012/
Fosberg FR, Stoddart DR, 1994. Flora of the Phoenix Islands, central Pacific. In: Atoll Research Bulletin, 393 60 pp.
Graveson R, 2012. Plants of Saint Lucia., http://www.saintlucianplants.com
Herrera K, Lorence D, Flynn T, Balick MJ, 2010. Checklist of the vascular plants of Pohnpei with local names and uses., Lawai, Hawaii, USA: National Tropical Botanical Garden. 146 pp.
Hokche O, Berry PE, Huber O, 2008. New catalogue of the vascular flora of Venezuela. (Nuevo Catálogo de la Flora Vascular de Venezuela)., Caracas, Venezuela, Fundación Instituto Botánico de Venezuela. 860 pp.
Madagascar Catalogue, 2012. Catalogue of the vascular plants of Madagascar. In: Missouri Botanical Garden, St Louis, USA: Missouri Botanical Garden and Madagascar Research and Conservation Program, Antananarivo. http://www.efloras.org/madagascar
Meyer JY, 2007. [English title not available]. (Rapport de mission sur l'Ile d'Uvea (Wallis & Futuna) du 6 au 17 Novembre 2007: Inventaire preliminaire de la flore vasculaire secondaire)., Papeete, Tahiti, Ministère de l'Education, l'Enseignement Supérieur et la Recherche. 39 pp. http://www.li-an.fr/jyves/Meyer_2007_Rapport_Plantes_Introduites_Wallis.pdf
Orchard AE, 1993. Flora of Australia. In: Oceanic islands 2, 50 Canberra, Australian Government Publishing Service.
PIER, 2013. Pacific Islands Ecosystems at Risk., Honolulu, Hawaii, USA: HEAR, University of Hawaii. http://www.hear.org/pier/index.html
Pruski JF, 2013. Asteraceae. In: Flora Mesoamericana, 5 (2) [ed. by Davidse G, Sousa Sánchez M, Knapp S, Chiang Cabrera F]. Mexico: Universidad Nacional Autónoma de México. http://www.tropicos.org/docs/meso/asteraceae.pdf
Smith AC, 1991. Flora Vitiensis nova: A new flora of Fiji., 5 Lawai, Kauai, Hawai`i, National Tropical Botanical Garden. 626 pp.
Space JC, Waterhouse BM, Newfield M, Bull C, 2004. Report to the Government of Niue and the United Nations Development Programme: Invasive Plant Species on Niue following Cyclone Heta., 76 pp. http://www.hear.org/pier/pdf/niue_report_20041217.pdf
Stone BC, 1970. The Flora of Guam. In: Micronesica, 6 1-659.
Thaman RR, Fosberg FR, Manner HI, Hassall DC, 1994. The flora of Nauru. In: Atoll Research Bulletin, 392 1-223.
USDA-ARS, 2013. Germplasm Resources Information Network (GRIN). Online Database. Beltsville, Maryland, USA: National Germplasm Resources Laboratory. https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysimple.aspx
Vibrans H, 2009. [English title not available]. (Malezas de México- Pennisetum purpureum)., http://www.conabio.gob.mx/malezasdemexico/poaceae/pennisetum-purpureum/fichas/ficha.htm
Wagner WL, Herbst DR, Sohmer SH, 1999. Manual of the flowering plants of Hawaii. Revised edition., Honolulu, Hawaii, USA: University of Hawaii Press/Bishop Museum Press. [Bernice P. Bishop Museum special publication].
Wagner WL, Herbst DR, Weitzman A, Lorence DH, 2013. Flora of Micronesia. Flora of Micronesia (online)., National Tropical Botanical Garden and the Smithsonian Institution. http://botany.si.edu/pacificislandbiodiversity/micronesia/index.htm
Whistler WA, 1983. The flora and vegetation of Swains Island. In: Atoll Research Bulletin, 262 25 pp.
Whistler WA, 1996. Botanical survey of Diego Garcia, Chagos Archipelago, British Indian Ocean Territory., Isle Botanica. 49 pp. http://www.zianet.com/tedmorris/dg/2005NRMP-Appendixe-botanicalsurvey.pdf
Links to Websites
Top of pageWebsite | URL | Comment |
---|---|---|
Catalogue of Seed Plants of the West Indies | http://botany.si.edu/antilles/WestIndies/catalog.htm | |
Global Compositae Checklist | http://compositae.landcareresearch.co.nz/Default.aspx | |
PIER | http://www.hear.org/pier/index.html | |
PROTA | http://www.prota4u.org/search.asp | |
Tropicos | http://www.tropicos.org/ |
Contributors
Top of page10/01/14 Original text by:
Julissa Rojas-Sandoval, Department of Botany-Smithsonian NMNH, Washington DC, USA
Pedro Acevedo-Rodríguez, Department of Botany-Smithsonian NMNH, Washington DC, USA
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