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Datasheet

Aspidiotus nerii (Oleander scale)

Summary

  • Last modified
  • 01 February 2017
  • Datasheet Type(s)
  • Pest
  • Natural Enemy
  • Preferred Scientific Name
  • Aspidiotus nerii
  • Preferred Common Name
  • Oleander scale
  • Taxonomic Tree
  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta

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Pictures

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PictureTitleCaptionCopyright
Adults and nymphs on citrus
TitleAdults and nymphs on citrus
Caption
CopyrightAgrEvo
Adults and nymphs on citrus
Adults and nymphs on citrusAgrEvo

Identity

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Preferred Scientific Name

  • Aspidiotus nerii Bouché, 1833

Preferred Common Name

  • Oleander scale

Other Scientific Names

  • Aspidiosus unipectinatus Ferris, 1941
  • Aspidiotus ? osmanthi Signoret, 1877
  • Aspidiotus affinis Targioni Tozzetti, 1868
  • Aspidiotus aloes Signoret, 1869
  • Aspidiotus atherospermae Maskell, 1879
  • Aspidiotus bouchei (Targioni Tozzetti) Targioni Tozzetti, 1868
  • Aspidiotus budlaei Maskell, 1879
  • Aspidiotus budleiae Signoret, 1869
  • Aspidiotus caldesii Targioni Tozzetti, 1868
  • Aspidiotus capparis Signoret, 1869
  • Aspidiotus carpodeti Maskell, 1885
  • Aspidiotus ceratoniae Signoret, 1869
  • Aspidiotus confusus Froggatt, 1914
  • Aspidiotus corynocarpi Colvée, 1881
  • Aspidiotus cycadicola (Boisduval) Signoret, 1869
  • Aspidiotus denticulatus Targioni Tozzetti, 1868
  • Aspidiotus dysoxyli Maskell, 1879
  • Aspidiotus epidendri Signoret, 1869
  • Aspidiotus ericae (Boisduval) Signoret, 1869
  • Aspidiotus fonsecai Giannotti, 1942
  • Aspidiotus genistae Westwood, 1840
  • Aspidiotus gnidii Signoret, 1869
  • Aspidiotus hederae Signoret, 1869
  • Aspidiotus hederae hederae Schmutterer, 1952
  • Aspidiotus hederae unipectinata Carimini, 1930
  • Aspidiotus hederae unisexualis Schmutterer, 1952
  • Aspidiotus hederae urenae Hall, 1923
  • Aspidiotus ilicis Signoret, 1869
  • Aspidiotus lentisci Signoret, 1877
  • Aspidiotus limonii Signoret, 1869
  • Aspidiotus myricinae Signoret, 1869
  • Aspidiotus myrsinae Signoret, 1877
  • Aspidiotus offinis Comstock, 1883
  • Aspidiotus oleae Colvée, 1880
  • Aspidiotus oleastri Colvée, 1882
  • Aspidiotus paranerii Gerson, in Gerson & Hazan, 1979
  • Aspidiotus rectangulatus (Lindinger) Ferris, 1941
  • Aspidiotus simillimus (Cockerell) Fernald, 1903
  • Aspidiotus sophorae Maskell, 1884
  • Aspidiotus tasmaniae Green, 1915
  • Aspidiotus transparens rectangulatus Lindinger, 1913
  • Aspidiotus transparens var. simillimus Cockerell, 1898
  • Aspidiotus transvaalensis Leonardi, 1914
  • Aspidiotus ulicis Signoret, 1869
  • Aspidiotus urenae (Hall) Ferris, 1941
  • Aspidiotus vagabundus Cockerell, 1899
  • Aspidiotus villosus Targioni Tozzetti, 1868
  • Aspidiotus viresciae Leonardi, 1920
  • Aspidiotus vriesciae Signoret, 1869
  • Chermes aloes Boisduval, 1867
  • Chermes cycadicola Boisduval, 1867
  • Chermes ericae Boisduval, 1867
  • Chermes genistae (Westwood) Ferris, 1941
  • Chermes hederae (Signoret) Ferris, 1937
  • Chermes nerii Boisduval, 1868
  • Chermes osmanthi Ferris, 1941
  • Coccus (Aspidiotus) palmarum Taschenberg, 1880
  • Coccus limonii Murray, 1871
  • Diaspis bouchei Targioni Tozzetti, 1867
  • Diaspis obliquum Costa, 1829
  • Octaspidiotus anthospermae Balachowsky, 1948
  • Octaspidiotus atherospermae (Maskell) MacGillivray, 1921

International Common Names

  • English: aucuba scale; ivy scale; lemon peel scale; white scale
  • Spanish: cochinilla blanca del olivo; piojo blanco; piojo blanco del naranjo; queresa del laurel
  • French: chermes du laurier-rose (m); cochenille du laurier-rose; cochenille du lierre
  • Russian: plushevaya shitovka
  • Portuguese: pinta-branca

Local Common Names

  • Chile: escama blanca de la hiedra
  • Denmark: palmeskjoldlus, gul
  • Finland: murattikilpikirva
  • Germany: Schildlaus, Oleander-
  • Israel: knimat hakisos
  • Italy: aspidioto dell'edera; bianca dei limoni; pidocchio bianco dei limoni
  • Japan: siro-marukaigaramusi
  • Mexico: escama hiedra
  • Netherlands: Kasschildluis
  • Norway: palmeskjoldlus
  • South Africa: oleanderdopluis
  • Sweden: palmskoeldlus, gul
  • Turkey: sarmasik kabuklu biti

EPPO code

  • ASPDNE (Aspidiotus nerii)

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta
  •                     Order: Hemiptera
  •                         Suborder: Sternorrhyncha
  •                             Superfamily: Coccoidea
  •                                 Family: Diaspididae
  •                                     Genus: Aspidiotus
  •                                         Species: Aspidiotus nerii

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

CountryDistributionLast ReportedOriginFirst ReportedInvasiveReferencesNotes

ASIA

ChinaPresentCIE, 1970
-ShandongPresentCIE, 1970
IranPresentNHM, 1973
IsraelPresentCIE, 1970
Japan
-HonshuPresentNHM, pre-1940
JordanPresentCIE, 1970
LebanonPresentCIE, 1970
Saudi ArabiaPresentCIE, 1970
SyriaPresentCIE, 1970
TurkeyPresentCIE, 1970; Ülgentürk et al., 2008

AFRICA

AlgeriaPresentCIE, 1970
Cape VerdePresentNHM, 1986
CongoPresentCIE, 1970
EgyptPresentCIE, 1970
EritreaPresentNHM, 1948
EthiopiaPresentCIE, 1970
KenyaPresentCIE, 1970
LibyaPresentCIE, 1970
MadagascarPresentCIE, 1970
MalawiPresentCIE, 1970
MoroccoPresentCIE, 1970
NigeriaPresentCIE, 1970
South AfricaPresentCIE, 1970
Spain
-Canary IslandsPresentCIE, 1970
TanzaniaPresentCIE, 1970
TunisiaPresentCIE, 1970
UgandaPresentCIE, 1970
ZimbabwePresentCIE, 1970

NORTH AMERICA

BermudaPresentNHM, 1925
MexicoPresentCIE, 1970
USAPresentCIE, 1970
-CaliforniaWidespreadShoemaker et al., 1979
-HawaiiPresentNHM, 1956

SOUTH AMERICA

ArgentinaPresentCIE, 1970
BoliviaPresentNHM, 1997
BrazilPresentCIE, 1970
-Espirito SantoPresentCulik et al., 2009
-Sao PauloPresentNHM, pre-1940
ChileWidespreadQuiroga et al., 1991; CIE, 1970
-Easter IslandPresentWilliams & Watson, 1988
ColombiaPresentCIE, 1970
EcuadorPresentNHM, 1982
PeruPresentCIE, 1970
UruguayPresentCIE, 1970

EUROPE

BelgiumPresentIntroducedMalumphy, 2016
BulgariaPresentCIE, 1970
CyprusPresentCIE, 1970
FrancePresentCIE, 1970
-CorsicaPresentCIE, 1970
GermanyPresentSchönfeld, 2015
GibraltarPresentCIE, 1970; NHM, 1930
GreecePresentAlexandrakis & Benassy, 1981; CIE, 1970
-CretePresentAlexandrakis & Benassy, 1981
IrelandPresentO'Connor et al., 2013
ItalyPresentCIE, 1970
-SicilyPresentRagusa et al., 2007
MaltaPresentCIE, 1970
PolandPresentGolan & Górska-Drabik, 2006; NHM, 1978
PortugalPresentCIE, 1970
-AzoresPresentCIE, 1970
-MadeiraPresentNHM, 1997
RomaniaPresentCIE, 1970
Russian FederationPresentCIE, 1970
SpainPresentCIE, 1970
-Balearic IslandsPresentCIE, 1970
UKPresentNHM, 1975
-England and WalesPresentNHM, 1990
Yugoslavia (former)PresentCIE, 1970

OCEANIA

AustraliaPresentCIE, 1970
-Lord Howe Is.PresentWilliams & Watson, 1988
-New South WalesPresentCIE, 1970
-QueenslandPresentCIE, 1970
-TasmaniaPresentNHM, 1989
New CaledoniaPresentWilliams & Watson, 1988; CIE, 1970
New ZealandPresentHill et al., 1993; CIE, 1970
Norfolk IslandPresentWilliams & Watson, 1988

Host plants/Plants Affected

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Plant nameFamilyContext
Actinidia chinensis (Chinese gooseberry)ActinidiaceaeMain
Albizia julibrissin (silk tree)FabaceaeMain
Ananas comosus (pineapple)BromeliaceaeOther
Asparagus setaceus (Asparagus fern)LiliaceaeOther
Asplenium nidus (bird's nest fern)AspleniaceaeOther
CeratoniaFabaceaeOther
CitrusRutaceaeMain
Cocos nucifera (coconut)ArecaceaeOther
Dianthus caryophyllus (carnation)CaryophyllaceaeOther
Diospyros (malabar ebony)EbenaceaeOther
Hedera helix (ivy)AraliaceaeOther
Ilex aquifolium (holly)AquifoliaceaeOther
Juniperus (junipers)CupressaceaeOther
KorthalsellaViscaceaeUnknown
Laurus nobilis (sweet bay)LauraceaeOther
MagnoliaMagnoliaceaeOther
Mangifera indica (mango)AnacardiaceaeOther
Melia azedarach (Chinaberry)MeliaceaeOther
Morus (mulberrytree)MoraceaeOther
Musa x paradisiaca (plantain)MusaceaeOther
Nerium (oleander)ApocynaceaeOther
Olea (olive)OleaceaeMain
Olea europaea subsp. europaea (European olive)OleaceaeMain
OleaceaeOleaceaeMain
Orchidaceae (orchids)OrchidaceaeOther
Pandanus (screw-pine)PandanaceaeOther
Phoenix (date palm)ArecaceaeOther
Pinus (pines)PinaceaeHabitat/association
Plumeria rubra var. acutifolia (Mexican frangipani)ApocynaceaeOther
Prunus persica (peach)RosaceaeOther
Pyrus communis (European pear)RosaceaeOther
Rosa (roses)RosaceaeOther
Simmondsia chinensis (jojoba)SimmondsiaceaeOther
Vitis vinifera (grapevine)VitaceaeOther

Growth Stages

Top of pageFlowering stage, Fruiting stage, Post-harvest, Seedling stage, Vegetative growing stage

List of Symptoms

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SignLife StagesType

Fruit

abnormal shape
discoloration
external feeding

Growing point

external feeding

Leaves

abnormal colours
abnormal forms
abnormal leaf fall
abnormal patterns
external feeding
wilting
yellowed or dead

Stems

dieback
external feeding
wilt

Whole plant

external feeding

Natural Enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Aleurodothrips fasciapennisPredator
Aphytis aonidiaeParasite
Aphytis chilensisParasiteAdults/Nymphs
Aphytis chrysomphaliParasiteAdults/Nymphs
Aphytis coheniParasiteAdults/Nymphs
Aphytis hispanicusParasite
Aphytis holoxanthusParasite
Aphytis lingnanensisParasiteAdults/Nymphs
Aphytis melinusParasiteAdults/NymphsChile; Crete; Sicily; SpainCitrus; olives
Aphytis paramaculicornisParasite
Chilocorus circumdatusPredatorAdults/Nymphs
Chilocorus infernalisPredatorAdults/Nymphs
Chilocorus kuwanaePredatorAdults/NymphsChina; HebeiJuniperus; Sabina
Chilocorus nigritaPredatorAdults/NymphsIsraelCitrus
Chilocorus renipustulatusPredatorAdults/Nymphs
Encarsia amiculaParasite
Encarsia aurantiiParasite
Encarsia citrinaParasiteAdults/Nymphs
Encarsia diaspidicolaParasite
Encarsia lounsburyiParasite
Habrolepis rouxiParasite
Hemisarcoptes coccophagusPredatorAdults/Nymphs
Hemisarcoptes malusParasite
Rhyzobius lophanthaePredatorAdults/Eggs/Larvae/Nymphs/Pupae
Rhyzobius pulchellusPredatorAdults/Nymphs
Signiphora flavopalliataParasite

References

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Alexandrakis V, Benassy C, 1981. Experiment in biological control on olive in Crete using Aphytis melinus De Bach (Hym. Aphelinidae), parasite of Aspidiotus nerii Bouche (Hom. Diaspididae). Acta Oecologica, Oecologia Applicata, 2(1):13-25.

Alexandrakis V, Neuenschwander P, 1980. The role of Aphytis chilensis (Hym.: Aphelinidae), a parasite of Aspidiotus nerii (Hom.: Diapsididae) on olive in Crete. Entomophaga, 25(1):61-71.

Badr SA, 2014. Insects and non insects species associated with pine needle trees in Alexandria Egypt. Journal of Entomology, 11(1):49-55. http://scialert.net/qredirect.php?doi=je.2014.49.55&linkid=pdf

Beardsley JW Jr, Gonzalez RH, 1975. The biology and ecology of armored scales [Diaspididae]. In: Smith RF, Mittler TE, Smith CN, ed. Annual Review of Entomology. Volume 20. Palo Alto, California, USA: Annual Reviews Inc., 47-73.

Ben-Dov Y, Matile-Ferrero D, 1999. Nomenclature of the Oleander scale (Hemiptera, Coccoidea, Diaspididae). Revue Française d'Entomologie, 21(1):5-8.

Booth RG, 1998. A review of the species resembling Chilocorus nigrita (Coleoptera: Coccinellidae): potential agents for biological control. Bulletin of Entomological Research, 88(4):361-367; 21 ref.

CIE, 1970. Distribution Maps of Pests, No. 268. Wallingford, UK: CAB International.

Claps LE, Terßn AL, 2001. Diaspididae (Hemiptera: Coccoidea) associated with citrus plants in Tucumßn, Argentina. Neotropical Entomology, 30(3):391-402; 38 ref.

Culik MP, Ventura JA, Martins Ddos S, 2009. Scale insects (Hemiptera: Coccidae) of pineapple in the State of Espírito Santo, Brazil. Acta Horticulturae [Proceedings of the Sixth International Pineapple Symposium, Joao Pessoa, Brazil, 18-23 November 2007.], No.822:215-218. http://www.actahort.org

Davidson RH, Lyon WF, 1987. Insect pests of farm, garden, and orchard. New York, USA: John Wiley & Sons, 8: 640 pp.

DeBach P, Rosen D, 1991. Biological control by natural enemies. Cambridge, UK: Cambridge University Press, 2:xiv + 440 pp.

Dreistadt SH, Clark JK, Flint ML, 1994. Pests of Landscape Trees and Shrubs: An Integrated Pest Management Guide. Oakland, California, USA: University of California, Agriculture and Natural Resources Publication 3359.

Dymock JJ, Holder PW, 1996. Nationwide survey of arthropods and molluscs on cut flowers in New Zealand. New Zealand Journal of Crop and Horticultural Science, 24(3):249-257.

Flint ML, 1990. Pests of the garden and small farm. A grower's guide to using less pesticide. Pests of the garden and small farm. A grower's guide to using less pesticide., x + 276 pp.

Golan K, Górska-Drabik E, 2006. The scale insects (Hemiptera, Coccinea) of ornamental plants in a greenhouse of the Maria Curie Skodowska University Botanical Garden in Lublin. Journal of Plant Protection Research, 46(4):347-352. http://www.ior.poznan.pl/Journal/

Grout TG, Moore SD, 2015. [English title not available]. In: Insects of Cultivated Plants and Natural Pastures in Southern Africa [ed. by Prinsloo, G. L., Uys, V. M.]. Hatfield, South Africa: Entomological Society of Southern Africa, 448-499.

Hare JD, Morgan DJW, 1997. Mass-priming Aphytis: behavioral improvement of insectary-reared biological control agents. Biological Control, 10(3):207-214; 29 ref.

Henderson RC, 2001. New synonymies for two armoured scale insects (Hemiptera: Coccoidea: Diaspididae) on Dysoxylum spectabile. New Zealand Entomologist, 24:89-90.

Henderson RC, Sultan A, Robertson AW, 2010. Scale insect fauna (Hemiptera: Sternorrhyncha: Coccoidea) of New Zealand's pygmy mistletoes (Korthalsella: Viscaceae) with description of three new species: Leucaspis albotecta, L. trilobata (Diaspididae) and Eriococcus korthalsellae (Eriococcidae). Zootaxa, 2644:1-24. http://www.mapress.com/zootaxa/2010/f/z02644p024f.pdf

Heu R, 1993. Distribution, Host Records of Agricultural Pests and Other Organisms of Hawaii. Hawaii, USA: State of Hawaii Deptartment of Agriculture, Survey Program, Pest Control Branch, Plant Industry Division.

Hill MG, Allan DJ, Henderson RC, Charles JG, 1993. Introduction of armoured scale predators and establishment of the predatory mite Hemisarcoptes coccophagus (Acari: Hemisarcoptidae) on latania scale, Hemiberlesia lataniae (Homoptera: Diaspididae) in kiwifruit shelter trees in New Zealand. Bulletin of Entomological Research, 83(3):369-376.

Honda JY, Luck RF, 1995. Scale morphology effects on feeding behaviour and biological control potential of Rhyzobius lophanthae (Coleoptera: Coccinellidae). Annals of the Entomological Society of America, 88(4):441-450; 34 ref.

Huffaker CB, Messenger PS, 1976. Theory and practice of biological control. Theory and practice of biological control., xxii + 788 pp.; [many fig., 240 X 170 mm].

Izraylevich S, Gerson U, 1993. Population dynamics of Hemisarcoptes coccophagus Meyer (Astigmata: Hemisarcoptidae) attacking three species of armored scale insects (Homoptera: Diaspididae). Experimental & Applied Acarology, 17(12):877-888.

Karaca I, Senal D, Colkesen T, ÷zg÷kce MS, 1999. Observations on the oleander scale, Aspidiotus nerii BouchT (Hemiptera: Diaspididae) and its natural enemies on blueleaf wattle in Adana Province, Turkey. Entomologica, 33:407-412; 5 ref.

Malumphy C, 2016. Six non-native species of scale insect (Hemiptera: Coccoidea) new for Belgium. Entomologist's Monthly Magazine, 152(4):245-251. http://www.pemberleybooks.com

O'Connor JP, Gertsson CA, Malumphy C, 2013. A review of the Irish scale insects (Hemiptera: Coccoidea). Irish Naturalists' Journal, 32(1):32-44. http://inj.biodiversityireland.ie/

Prinsloo GL Uys VM, 2015. Insects of Cultivated Plants and Natural Pastures in Southern Africa. Hatfield, South Africa: Entomological Society of Southern Africa.

Quiroga D, Arretz P, Araya JE, 1991. Sucking insects damaging jojoba, Simmondsia chinesis (Link) Schneider, and their natural enemies, in the North Central and Central regions of Chile. Crop Protection, 10(6):469-472.

Ragusa S, Tsolakis H, Caleca V, Agrò A, Pinto Mlo, 2007. Observations on citrus groves acarofauna and control techniques of phytophagous mites, Phyllocnistis citrella and Aspidiotus nerii. (Osservazioni sull'acarocenosi dell'agrumeto e tecniche di controllo di acari fitofagi, di Phyllocnistis citrella e di Aspidiotus nerii.) Italus Hortus [Atti del Convegno Risultati finali progetto MiPAAF "Ricerche e sperimentazioni nel settore dell'agrumicoltura italiana" (Piano Agrumicolo Nazionale), Acireale (CT), Italy, 30 maggio-1 giugno 2007.], 14(6):108-109. http://www.soihs.it

Rocha KCG, Silva RAda, Michelotto MD, Busoli AC, 2006. Biological, morphological and behavioral aspects of Aspidiotus nerii Bouché, 1833 (Hemiptera: Diaspididae). (Aspectos biológicos, morfológicos e comportamentais de Aspidiotus nerii Bouché, 1833 (Hemiptera: Diaspididae).) Ciência Rural, 36(2):363-368. http://www.ufsm.br/ccr/revista

Schönfeld U, 2015. Coccoidea species in Brandenburg. (Cocciden-arten in Brandenburg.) Journal für Kulturpflanzen, 67(10):337-341. http://www.journal-kulturpflanzen.de

Shoemaker CA, Huffaker CB, Kennett CE, 1979. A systems approach to the integrated management of a complex of olive pests. Environmental Entomology, 8(1):182-189.

Signoret V, 1868. Essai sur les cochenilles (HomoptFres-Coccides), 2e partie. Annales de la Société entomologique de France (4)8:503-528.

Signoret V, 1869. Essai sur les cochenilles ou gallinsectes (Homoptères-Coccides), 3e partie. Annales de la Société entomologique de France, 4(9):97-104.

Stathas GJ, Bouras SL, Eliopoulos PA, Emmanouel NG, 2005. Control of diaspidid scales on olive trees by releasing coccinellid predators. Bulletin OILB/SROP [Proceedings of the European meeting of the IOBC/WPRS Working Group "Integrated Protection of Olive Crops", Chania, Greece, 29-31 May 2003.], 28(9):157-166.

Swan LA, 1964. Beneficial Insects. New York, USA: Harper and Row.

Therios I, 2008. Pests and diseases. In: Olives [ed. by Therios, I.]. Wallingford, UK: CABI, 335-352. http://www.cabi.org/CABeBooks/default.aspx?site=107&page=45&LoadModule=PDFHier&BookID=440

Ülgentürk S, Sahin Ö, Kaydan MB, 2008. Coccoidea (Hemiptera) species on park plants in urban areas of Istanbul province. (Istanbul ili yesil alan bitkilerinde bulunan Coccoidea (Hemiptera) türleri.) Bitki Koruma Bülteni, 48(1):1-18. http://www.ziraimucadele.gov.tr

VanDriesche R, Idione K, Rose M, Bryan M, 1998. Evaluation of the effectiveness of Chilocorus kuwanae (Coleoptera: Coccinellidae) in suppressing the euonymus scale (Homoptera: Diaspididae). Biological Control, 12: 56-65.

Williams DJ, Watson GW, 1988. The Scale Insects of the Tropical South Pacific Region. Part 1. The Armoured Scales (Diaspididae). Wallingford, UK: CAB International.

Distribution Maps

Top of page
Distribution map Argentina: Present
CIE, 1970Australia: Present
CIE, 1970Australia
See regional map for distribution within the countryAustralia
See regional map for distribution within the countryAustralia
See regional map for distribution within the countryAustralia
See regional map for distribution within the countryBelgium: Present, introduced
Malumphy, 2016Bulgaria: Present
CIE, 1970Bermuda: PresentBolivia: PresentBrazil: Present
CIE, 1970Brazil
See regional map for distribution within the countryBrazil
See regional map for distribution within the countryCongo: Present
CIE, 1970Chile: Widespread
Quiroga et al., 1991; CIE, 1970Chile
See regional map for distribution within the countryChina: Present
CIE, 1970China: Present
CIE, 1970China
See regional map for distribution within the countryColombia: Present
CIE, 1970Colombia: Present
CIE, 1970Cape Verde: PresentCyprus: Present
CIE, 1970Cyprus: Present
CIE, 1970Germany: Present
Schönfeld, 2015Algeria: Present
CIE, 1970Algeria: Present
CIE, 1970Ecuador: PresentEgypt: Present
CIE, 1970Eritrea: PresentSpain: Present
CIE, 1970Spain: Present
CIE, 1970Spain
See regional map for distribution within the countrySpain
See regional map for distribution within the countryEthiopia: Present
CIE, 1970France: Present
CIE, 1970France
See regional map for distribution within the countryUK: PresentUK
See regional map for distribution within the countryGibraltar: Present
CIE, 1970Greece: Present
Alexandrakis & Benassy, 1981; CIE, 1970Greece: Present
Alexandrakis & Benassy, 1981; CIE, 1970Greece
See regional map for distribution within the countryGreece
See regional map for distribution within the countryIreland: Present
O'Connor et al., 2013Israel: Present
CIE, 1970Israel: Present
CIE, 1970Iran: PresentIran: PresentIran: PresentItaly: Present
CIE, 1970Italy
See regional map for distribution within the countryItaly
See regional map for distribution within the countryJordan: Present
CIE, 1970Jordan: Present
CIE, 1970Japan
See regional map for distribution within the countryKenya: Present
CIE, 1970Lebanon: Present
CIE, 1970Lebanon: Present
CIE, 1970Lebanon: Present
CIE, 1970Libya: Present
CIE, 1970Morocco: Present
CIE, 1970Morocco: Present
CIE, 1970Madagascar: Present
CIE, 1970Malta: Present
CIE, 1970Malta: Present
CIE, 1970Malawi: Present
CIE, 1970Mexico: Present
CIE, 1970Mexico: Present
CIE, 1970New Caledonia: Present
Williams & Watson, 1988; CIE, 1970Norfolk Island: Present
Williams & Watson, 1988Nigeria: Present
CIE, 1970New Zealand: Present
Hill et al., 1993; CIE, 1970Peru: Present
CIE, 1970Poland: Present
Golan & Górska-Drabik, 2006Portugal: Present
CIE, 1970Portugal
See regional map for distribution within the countryPortugal
See regional map for distribution within the countryPortugal
See regional map for distribution within the countryRomania: Present
CIE, 1970Russian Federation: Present
CIE, 1970Russian Federation: Present
CIE, 1970Saudi Arabia: Present
CIE, 1970Saudi Arabia: Present
CIE, 1970Syria: Present
CIE, 1970Syria: Present
CIE, 1970Syria: Present
CIE, 1970Tunisia: Present
CIE, 1970Tunisia: Present
CIE, 1970Turkey: Present
CIE, 1970; Ülgentürk et al., 2008Turkey: Present
CIE, 1970; Ülgentürk et al., 2008Turkey: Present
CIE, 1970; Ülgentürk et al., 2008Tanzania: Present
CIE, 1970Uganda: Present
CIE, 1970USA: Present
CIE, 1970USA: Present
CIE, 1970USA
See regional map for distribution within the countryUSA
See regional map for distribution within the countryUruguay: Present
CIE, 1970Yugoslavia (former): Present
CIE, 1970South Africa: Present
CIE, 1970Zimbabwe: Present
CIE, 1970
  • = Present, no further details
  • = Evidence of pathogen
  • = Widespread
  • = Last reported
  • = Localised
  • = Presence unconfirmed
  • = Confined and subject to quarantine
  • = See regional map for distribution within the country
  • = Occasional or few reports
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Distribution map (asia) China: Present
CIE, 1970Shandong: Present
CIE, 1970Israel: Present
CIE, 1970Iran: PresentJordan: Present
CIE, 1970Honshu: PresentLebanon: Present
CIE, 1970Russian Federation: Present
CIE, 1970Saudi Arabia: Present
CIE, 1970Syria: Present
CIE, 1970Turkey: Present
CIE, 1970; Ülgentürk et al., 2008
Distribution map (europe) Belgium: Present, introduced
Malumphy, 2016Bulgaria: Present
CIE, 1970Cyprus: Present
CIE, 1970Germany: Present
Schönfeld, 2015Algeria: Present
CIE, 1970Spain: Present
CIE, 1970Balearic Islands: Present
CIE, 1970France: Present
CIE, 1970Corsica: Present
CIE, 1970UK: PresentEngland and Wales: PresentGibraltar: Present
CIE, 1970Greece: Present
Alexandrakis & Benassy, 1981; CIE, 1970Crete: Present
Alexandrakis & Benassy, 1981Ireland: Present
O'Connor et al., 2013Iran: PresentItaly: Present
CIE, 1970Sicily: Present
Ragusa et al., 2007Lebanon: Present
CIE, 1970Morocco: Present
CIE, 1970Malta: Present
CIE, 1970Poland: Present
Golan & Górska-Drabik, 2006Portugal: Present
CIE, 1970Azores: Present
CIE, 1970Madeira: PresentRomania: Present
CIE, 1970Russian Federation: Present
CIE, 1970Syria: Present
CIE, 1970Tunisia: Present
CIE, 1970Turkey: Present
CIE, 1970; Ülgentürk et al., 2008Yugoslavia (former): Present
CIE, 1970
Distribution map (africa) Congo: Present
CIE, 1970Cape Verde: PresentCyprus: Present
CIE, 1970Algeria: Present
CIE, 1970Egypt: Present
CIE, 1970Eritrea: PresentSpain: Present
CIE, 1970Canary Islands: Present
CIE, 1970Ethiopia: Present
CIE, 1970Greece: Present
Alexandrakis & Benassy, 1981; CIE, 1970Crete: Present
Alexandrakis & Benassy, 1981Israel: Present
CIE, 1970Iran: PresentSicily: Present
Ragusa et al., 2007Jordan: Present
CIE, 1970Kenya: Present
CIE, 1970Lebanon: Present
CIE, 1970Libya: Present
CIE, 1970Morocco: Present
CIE, 1970Madagascar: Present
CIE, 1970Malta: Present
CIE, 1970Malawi: Present
CIE, 1970Nigeria: Present
CIE, 1970Madeira: PresentSaudi Arabia: Present
CIE, 1970Syria: Present
CIE, 1970Tunisia: Present
CIE, 1970Turkey: Present
CIE, 1970; Ülgentürk et al., 2008Tanzania: Present
CIE, 1970Uganda: Present
CIE, 1970South Africa: Present
CIE, 1970Zimbabwe: Present
CIE, 1970
Distribution map (north america) Bermuda: PresentMexico: Present
CIE, 1970USA: Present
CIE, 1970California: Widespread
Shoemaker et al., 1979Hawaii: Present
Distribution map (central america) Colombia: Present
CIE, 1970Mexico: Present
CIE, 1970USA: Present
CIE, 1970
Distribution map (south america) Argentina: Present
CIE, 1970Bolivia: PresentBrazil: Present
CIE, 1970Espirito Santo: Present
Culik et al., 2009Sao Paulo: PresentChile: Widespread
Quiroga et al., 1991; CIE, 1970Easter Island: Present
Williams & Watson, 1988Colombia: Present
CIE, 1970Ecuador: PresentPeru: Present
CIE, 1970Uruguay: Present
CIE, 1970
Distribution map (pacific) Australia: Present
CIE, 1970Lord Howe Is.: Present
Williams & Watson, 1988New South Wales: Present
CIE, 1970Queensland: Present
CIE, 1970Tasmania: PresentChina: Present
CIE, 1970New Caledonia: Present
Williams & Watson, 1988; CIE, 1970Norfolk Island: Present
Williams & Watson, 1988New Zealand: Present
Hill et al., 1993; CIE, 1970