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Datasheet

Planococcus lilacinus (cacao mealybug)

Summary

  • Last modified
  • 20 January 2015
  • Datasheet Type(s)
  • Pest
  • Preferred Scientific Name
  • Planococcus lilacinus
  • Preferred Common Name
  • cacao mealybug
  • Taxonomic Tree
  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta

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Pictures

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PictureTitleCaptionCopyright
P. lilacinus - slide mounted adult female.
TitleAdult female - line drawing
CaptionP. lilacinus - slide mounted adult female.
CopyrightBeatrice Padi (modified from Cox, 1989)
P. lilacinus - slide mounted adult female.
Adult female - line drawingP. lilacinus - slide mounted adult female.Beatrice Padi (modified from Cox, 1989)
P. lilacinus (2.7mm) on cocoa pod.
TitleP. lilacinus on cocoa
CaptionP. lilacinus (2.7mm) on cocoa pod.
CopyrightPeter A.C. Ooi/Tropical Press Sdn Bnd
P. lilacinus (2.7mm) on cocoa pod.
P. lilacinus on cocoaP. lilacinus (2.7mm) on cocoa pod.Peter A.C. Ooi/Tropical Press Sdn Bnd

Identity

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Preferred Scientific Name

  • Planococcus lilacinus Cock

Preferred Common Name

  • cacao mealybug

Other Scientific Names

  • Dactylopius coffeae Newstead
  • Dactylopius crotonis Green
  • Planococcus crotonis (Green)
  • Planococcus deceptor
  • Planococcus tayabanus (Cockerell)
  • Pseudococcus coffeae (Newstead)
  • Pseudococcus crotonis (Green)
  • Pseudococcus deceptor Betrem
  • Pseudococcus lilacinus Cockerell
  • Pseudococcus tayabanus Cockerell
  • Tylococcus mauritiensis Mamet

International Common Names

  • English: coffee mealybug

Local Common Names

  • Germany: Asiatische Kaffee-Schmierlaus; Kaffeewurzel-Schmierlaus

EPPO code

  • PLANLI (Planococcus lilacinus)

Taxonomic Tree

Top of page
  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta
  •                     Order: Hemiptera
  •                         Suborder: Sternorrhyncha
  •                             Superfamily: Coccoidea
  •                                 Family: Pseudococcidae
  •                                     Genus: Planococcus
  •                                         Species: Planococcus lilacinus

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

CountryDistributionLast ReportedOriginFirst ReportedInvasiveReferencesNotes

ASIA

BangladeshPresentCox, 1989; Ben-Dov, 1994; EPPO, 2014
Brunei DarussalamPresentWilliams, 1982; EPPO, 2014
CambodiaPresentBen-Dov, 1994; Takahashi, 1942; EPPO, 2014
Cocos IslandsPresentCox, 1989; EPPO, 2014
IndiaRestricted distributionEPPO, 2014
-Andaman and Nicobar IslandsPresentIIE, 1995; EPPO, 2014
-BiharPresentIIE, 1995; EPPO, 2014
-DelhiPresentButani, 1978; IIE, 1995; EPPO, 2014
-GujaratPresentIIE, 1995; EPPO, 2014
-KarnatakaPresentMani & Krishnamoorthy, 1990; Reddy et al., 1992; EPPO, 2014
-KeralaPresentPillai, 1987; Kumar & Prakasan, 1992; EPPO, 2014
-MaharashtraPresentIIE, 1995; EPPO, 2014
-OdishaPresentIIE, 1995; EPPO, 2014
-Tamil NaduPresentDhandapani et al., 1992; IIE, 1995; EPPO, 2014
IndonesiaPresentEPPO, 2014
-Irian JayaPresentWilliams & Watson, 1988; EPPO, 2014
-JavaPresentLe, 1943; Cox, 1989; IIE, 1995; EPPO, 2014
-KalimantanPresentBen-Dov, 1994; EPPO, 2014
-Nusa TenggaraPresentEPPO, 2014
-SulawesiPresentIIE, 1995; EPPO, 2014
JapanRestricted distributionEPPO, 2014
-Ryukyu ArchipelagoPresentIIE, 1995; EPPO, 2014
LaosPresentIIE, 1995; EPPO, 2014
MalaysiaRestricted distributionEPPO, 2014
-Peninsular MalaysiaRestricted distributionIIE, 1995; EPPO, 2014
-SabahPresentIIE, 1995; EPPO, 2014
-SarawakPresentWilliams, 1982; IIE, 1995; EPPO, 2014
MaldivesPresentIIE, 1995; EPPO, 2014
MyanmarPresentWilliams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014
PhilippinesPresentCox, 1989; Ben-Dov, 1994; EPPO, 2014
Sri LankaPresentBen-Dov, 1994; EPPO, 2014; Fernando & Karagaratram, 1987
TaiwanPresentCox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014
ThailandPresentBen-Dov, 1994; EPPO, 2014; Takashi, 1942
VietnamPresentCox, 1989; Ben-Dov, 1994; EPPO, 2014
YemenPresentCox, 1989; Ben-Dov, 1994; IIE, 1995; EPPO, 2014

AFRICA

AfricaPresentWilliams et al., 2001
ComorosPresentBen-Dov, 1994; EPPO, 2014
KenyaPresent, few occurrencesBen-Dov, 1994; IIE, 1995; EPPO, 2014
MadagascarPresentCox, 1989; Ben-Dov, 1994; EPPO, 2014
MauritiusPresentCox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014
RéunionPresentWilliams et al., 2001
SeychellesPresentWilliams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014

CENTRAL AMERICA AND CARIBBEAN

Dominican RepublicPresentWilliams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014
El SalvadorPresentWilliams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014
HaitiPresentWilliams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014

SOUTH AMERICA

GuyanaPresentWilliams, 1982; Williams & Granara, 1992; EPPO, 2014

OCEANIA

GuamPresentEPPO, 2014
Micronesia, Federated states ofPresentEPPO, 2014
Northern Mariana IslandsPresentBeardsley, 1966
Papua New GuineaPresentWilliams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014

Host plants/Plants Affected

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Plant nameFamilyContext
Annona muricata (soursop)AnnonaceaeMain
Annona squamosa (sugar apple)AnnonaceaeOther
Bambusa vulgaris (common bamboo)PoaceaeOther
Bauhinia (camel's foot)FabaceaeMain
Brassica oleracea var. botrytis (cauliflower)BrassicaceaeOther
Ceiba pentandra (kapok)BombacaceaeMain
CitrusRutaceaeMain
Cocos nucifera (coconut)ArecaceaeOther
Coffea (coffee)RubiaceaeMain
Coffea arabica (arabica coffee)RubiaceaeMain
Coffea canephora (robusta coffee)RubiaceaeMain
Dioscorea (yam)DioscoreaceaeOther
Mangifera indica (mango)AnacardiaceaeOther
Psidium guajava (guava)MyrtaceaeMain
Punica granatum (pomegranate)PunicaceaeOther
Tamarindus indica (Indian tamarind)FabaceaeOther
Theobroma cacao (cocoa)SterculiaceaeMain
Vitis (grape)VitaceaeOther

Growth Stages

Top of pageFlowering stage, Fruiting stage, Seedling stage, Vegetative growing stage

List of Symptoms

Top of page
SignLife StagesType

Fruit

abnormal shape
external feeding
premature drop

Growing point

dieback

Inflorescence

wilt

Leaves

honeydew or sooty mould
honeydew or sooty mould
necrotic areas

Stems

external feeding
honeydew or sooty mould
necrosis

Whole plant

wilt

Natural Enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Aenasius lepelleyiParasiteAdults/Nymphs
Aenasius subbaraoiParasiteAdults/Nymphs
AnagyrusParasiteAdults/Nymphs
Anagyrus pseudococciParasite
Brumoides suturalisPredatorAdults/Nymphs
Coccodiplosis smithiPredatorAdults/Nymphs
Cryptolaemus montrouzieriPredatorAdults/NymphsIndia
Eublemma scitulaPredatorAdults/Nymphs
Mallada boninensisPredatorAdults/Nymphs
Miletus boisduvaliPredatorAdults/Nymphs
Promuscidea unfasciativentrisParasiteAdults/Nymphs
Pseudaphycus orientalisParasiteAdults/Nymphs
Pseudoscymnus pallidicollisPredatorAdults/Nymphs
Scymnus apiciflavusPredatorAdults/Nymphs
Scymnus roepkeiPredatorAdults/Nymphs
Spalgis epeusPredatorAdults/Nymphs
Tetracnemoidea indicaParasite
Triommata coccidivoraPredatorAdults/Nymphs

References

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1978. Thirtyfirst annual report 1977-78. Report, Coffee Board Research Department, India, [2+] 53 pp.

Avasti RK, Saafe SA, 1987. Indian Pseudococcidae (Homoptera: Coccoidea). Indian Journal of Systematic Entomology, 4:1-54.

Beardsley JW, 1966. Insects of Micronesia, 6(7):434-435.

Ben-Dov Y, 1994. A systematic catalogue of the mealybugs of the world (Insecta: Homoptera: Coccoidea: Pseudococcidae and Putoidae) with data on geographical distribution, host plants, biology and economic importance. Andover, UK; Intercept Limited, 686 pp.

Bhat PK, Shamanna HV, 1972. Some new collateral hosts of Planococcus lilacinus from South India. Journal of Coffee Research, 2(2):27

Brough EJ, 1980. Citrus entomology in Papua New Guinea. News Bulletin, Entomological Society of Queensland, 14:43-47.

Butani DK, 1976. Insect pests of fruit crops and their control - custard apple. Pesticides, 10(5):27-28

Butani DK, 1978. Insect pests of tamarind and their control. Pesticides, 12(11):34-41

Chacko MJ, Bhat PK, Rao LVAnanda, Deepak Singh MB, Ramanarayan EP, Sreedharan K, 1978. The use of the ladybird beetle, Cryptolpmus montrouzieri for the control of coffee mealybugs. Journal of Coffee Research, 8(1):14-19

Chacko MJ, Bhat PKrishnamoorthy, Ramanarayan EP, 1977. New records of Coccoidea with notes on natural enemies of Planococcus spp. on coffee in India. Journal of Coffee Research, 7(3):69-71

Chacko MJ, Sreedharan K, 1981. Predation of Mallada boninensis on Ferrisia virgata, Planococcus citri and P. lilacinus. Journal of Biological Control, 4(2):122-123.

Cox JM, 1989. The mealybug genus Planococcus (Homoptera: Pseudococcidae). Bulletin of the British Museum (Natural History), Entomology, 58(1):1-78

Dayan MP, Baltazar EM, 1990. Survey, identification and pathogenicity of pests and diseases of bamboo in the Philippines. Proceedings of the first National Bamboo Symposium-Workshop, February 27-March 01, 1989. Sylvatrop, 13(1-2):61-77.

Dhandapani N, Gopalan M, Sundarababu PC, 1992. Evaluation of insecticides for the control of mealy bugs, (Planococcus lilacinus, Ckll.) in jasmine. Madras Agricultural Journal, 79(1):54-55

Entwistle PF, 1972. Pests of cocoa. London, UK: Longman, 779 pp.

EPPO, 2014. PQR database. Paris, France: European and Mediterranean Plant Protection Organization. http://www.eppo.int/DATABASES/pqr/pqr.htm

Fernando LCP, Kanagaratnam P, 1987. New records of some pests of the coconut inflorescence and developing fruit and their natural enemies in Sri Lanka. COCOS, 5:39-42.

Ferris GF, 1950. Atlas of the Scale Insects of North America. Series V. The Pseudococeidae (Part I). Stanford Univ. Pr., California; London, G. Cumberlege, Oxford Univ. Pr., vii+ 278 pp.

Graham MK, 1991. Biological control of Helopeltis spp. in mature cocoa by the black ant (Dolichoderus bituberculatus) and the cocoa mealybug (Planococcus lilacinus). Planter, 67(788):543-546

IIE, 1995. Distribution Maps of Pests, Series A, No. 101. Wallingford, UK: CAB International.

Kumar MG, Bhat PK, Ramaiah PK, 1989. Potential role of kerosene and neem derivatives in integrated management of mealybugs on coffee. Journal of Coffee Research, 19(1):17-29

Kumar PKV, Prakasan CB, 1992. Soil application of systemic insecticides for mealybug control. Journal of Coffee Research, 22(1):65-68

Le Pelley RH, 1943. An Oriental mealybug (Pseudococcus lilacinus Ckll.) (Hemiptera) and its insect enemies. Transactions of the Royal Entomological Society of London, 93(1):73-93.

Mani M, 1995. Comparative development, progeny production and sex ratio of the exotic parasitoid Leptomastix dactylopii Howard (Hym., Encyrtidae) on Planococcus lilacinus and P. citri (Homop., Pseudococcidae). Entomon, 20(1):23-26

Mani M, 1995. Studies on the natural enemies of oriental mealybug, Planococcus lilacinus (Ckll.) (Homoptera: Pseudococcidae) in India. Journal of Entomological Research, 19(1):61-70

Mani M, Krishnamoorthy A, 1990. Predation of Mallada boninensis on Ferrisia virgata, Planococcus citri and P. lilacinus. Journal of Biological Control, 4(2):122-123

McKenzie HL, 1967. Mealybugs of California, with taxonomy, biology, and control of North American species (Homoptera: Coccoidea: Pseudococcidae). viii+526 pp.

Nagarkatti S, Singh SP, Jayanth KP, Bhummannavar BS, 1992. Introduction and establishment of Leptomastix dactylopii How. against Planococcus spp. in India. Indian Journal of Plant Protection, 20(1):102-104

Noyes JS, Hayat M, 1994. Oriental mealybug parasitoids of the Anagyrini (Hymenoptera: Encyrtidae). Wallingford, UK; CAB International, viii + 554 pp.

Pelley RH le, 1968. Pests of Coffee. London and Harlow, UK: Longmans, Green and Co Ltd.

Pillai GB, 1987. Integrated pest management in plantation crops. Journal of Coffee Research, 17(1):150-153

Prakasan CB, 1987. Biological control of coffee pests. Journal of Coffee Research, 17(1):114-117

Reddy KB, Prakasan CB, Bhat PK, Kumar AC, 1992. Establishment of Leptomastix dactylopii How. (Hym.: Encyrtidae) in Karnataka for control of Planococcus citri (Risso) (Hom.: Pseudococcidae) of coffee. Journal of Coffee Research, 22(1):37-44

Sekhar PS, 1964. Pests of coffee. In: Entomology in India. Indian Journal of Entomology, 99-109.

Shukla RP, Tandon PL, 1984. Indiainsect pests on custard apple. Plant Protection Bulletin, FAO, 32(1):31

Szent-Ivany JJH, Catley A, 1960. Host plant and distribution records of some insects in New Guinea and adjacent islands. Pacific Insects, 2:255-261.

Takahashi R, 1942. Report Government Agricultural Research Institute, Taiwan, 81:10.

Tandon PL, Verghese A, 1987. New insect pests of certain fruit crops. Indian Journal of Horticulture, 44(1-2):121-122

van der Goot P, 1917. De zwarte cacao-mier (Dolichoderus bituberculatus, Mayr.) en haar Beteekenis voor de cacaocultuur op Java, Meded. Proefst. Midden-Java, Salatiga, 25:142pp.

Watson GW, Cox JM, 1990. Identity of the African coffee root mealybug, with descriptions of two new species of Planococcus (Homoptera: Pseudococcidae). Bulletin of Entomological Research, 80(1):99-105

Williams DJ, 1982. The distribution of the mealybug genus Planococcus (Hemiptera: Pseudococcidae) in Melanesia, Polynesia and Kiribati. Bulletin of Entomological Research, 72(3):441-455

Williams DJ, Granara de Willink MC, 1992. Mealybugs of Central and South America. Wallingford, UK: CAB International.

Williams DJ, Matile-Ferrero D, Martin JH, 2001. The mealybug Planococcus lilacinus (Cockerell) in Africa (Hemiptera, Coccoidea, Pseudococcidae). Bulletin de la Société Entomologique de France, 106(3):259-260.

Williams DJ, Watson GW, 1988. Scale insects of the tropical South Pacific region. Part 2. Mealybugs (Pseudococcidae). Wallingford, Oxon, UK; CAB International, 260 pp.

Distribution Maps

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Distribution map Africa: Present
Williams et al., 2001Bangladesh: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Brunei Darussalam: Present
Williams, 1982; EPPO, 2014Cocos Islands: Present
Cox, 1989; EPPO, 2014Dominican Republic: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014Dominican Republic: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014Micronesia, Federated states of: Present
EPPO, 2014Guam: Present
EPPO, 2014Guyana: Present
Williams, 1982; Williams & Granara, 1992; EPPO, 2014Guyana: Present
Williams, 1982; Williams & Granara, 1992; EPPO, 2014Haiti: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014Haiti: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014Indonesia: Present
EPPO, 2014Indonesia: Present
EPPO, 2014Indonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndia: Restricted distribution
EPPO, 2014India
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryJapan: Restricted distribution
EPPO, 2014Japan
See regional map for distribution within the countryKenya: Present, few occurrences
Ben-Dov, 1994; IIE, 1995; EPPO, 2014Cambodia: Present
Ben-Dov, 1994; Takahashi, 1942; EPPO, 2014Comoros: Present
Ben-Dov, 1994; EPPO, 2014Laos: Present
IIE, 1995; EPPO, 2014Sri Lanka: Present
Ben-Dov, 1994; EPPO, 2014Madagascar: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Myanmar: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014Northern Mariana Islands: Present
Beardsley, 1966Mauritius: Present
Cox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014Maldives: Present
IIE, 1995; EPPO, 2014Malaysia: Restricted distribution
EPPO, 2014Malaysia
See regional map for distribution within the countryMalaysia
See regional map for distribution within the countryMalaysia
See regional map for distribution within the countryPapua New Guinea: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014Papua New Guinea: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014Philippines: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Philippines: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Réunion: Present
Williams et al., 2001Seychelles: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014El Salvador: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014El Salvador: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014Thailand: Present
Ben-Dov, 1994; EPPO, 2014Taiwan: Present
Cox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014Taiwan: Present
Cox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014Vietnam: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Yemen: Present
Cox, 1989; Ben-Dov, 1994; IIE, 1995; EPPO, 2014Yemen: Present
Cox, 1989; Ben-Dov, 1994; IIE, 1995; EPPO, 2014
  • = Present, no further details
  • = Evidence of pathogen
  • = Widespread
  • = Last reported
  • = Localised
  • = Presence unconfirmed
  • = Confined and subject to quarantine
  • = See regional map for distribution within the country
  • = Occasional or few reports
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Distribution map (asia) Bangladesh: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Brunei Darussalam: Present
Williams, 1982; EPPO, 2014Cocos Islands: Present
Cox, 1989; EPPO, 2014Indonesia: Present
EPPO, 2014Irian Jaya: Present
Williams & Watson, 1988; EPPO, 2014Java: Present
Le, 1943; Cox, 1989; IIE, 1995; EPPO, 2014Kalimantan: Present
Ben-Dov, 1994; EPPO, 2014Sulawesi: Present
IIE, 1995; EPPO, 2014India: Restricted distribution
EPPO, 2014Andaman and Nicobar Islands: Present
IIE, 1995; EPPO, 2014Bihar: Present
IIE, 1995; EPPO, 2014Delhi: Present
Butani, 1978; IIE, 1995; EPPO, 2014Gujarat: Present
IIE, 1995; EPPO, 2014Kerala: Present
Pillai, 1987; Kumar & Prakasan, 1992; EPPO, 2014Karnataka: Present
Mani & Krishnamoorthy, 1990; Reddy et al., 1992; EPPO, 2014Maharashtra: Present
IIE, 1995; EPPO, 2014Odisha: Present
IIE, 1995; EPPO, 2014Tamil Nadu: Present
Dhandapani et al., 1992; IIE, 1995; EPPO, 2014Japan: Restricted distribution
EPPO, 2014Ryukyu Archipelago: Present
IIE, 1995; EPPO, 2014Cambodia: Present
Ben-Dov, 1994; Takahashi, 1942; EPPO, 2014Laos: Present
IIE, 1995; EPPO, 2014Sri Lanka: Present
Ben-Dov, 1994; EPPO, 2014Myanmar: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014Maldives: Present
IIE, 1995; EPPO, 2014Malaysia: Restricted distribution
EPPO, 2014Peninsular Malaysia: Restricted distribution
IIE, 1995; EPPO, 2014Sabah: Present
IIE, 1995; EPPO, 2014Sarawak: Present
Williams, 1982; IIE, 1995; EPPO, 2014Papua New Guinea: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014Philippines: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Thailand: Present
Ben-Dov, 1994; EPPO, 2014Taiwan: Present
Cox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014Vietnam: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Yemen: Present
Cox, 1989; Ben-Dov, 1994; IIE, 1995; EPPO, 2014
Distribution map (europe)
Distribution map (africa) Africa: Present
Williams et al., 2001Kenya: Present, few occurrences
Ben-Dov, 1994; IIE, 1995; EPPO, 2014Comoros: Present
Ben-Dov, 1994; EPPO, 2014Madagascar: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Mauritius: Present
Cox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014Réunion: Present
Williams et al., 2001Seychelles: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014Yemen: Present
Cox, 1989; Ben-Dov, 1994; IIE, 1995; EPPO, 2014
Distribution map (north america) Dominican Republic: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014Haiti: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014El Salvador: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014
Distribution map (central america) Dominican Republic: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014Guyana: Present
Williams, 1982; Williams & Granara, 1992; EPPO, 2014Haiti: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014El Salvador: Present
Williams & Granara, 1992; Ben-Dov, 1994; EPPO, 2014
Distribution map (south america) Guyana: Present
Williams, 1982; Williams & Granara, 1992; EPPO, 2014
Distribution map (pacific) Micronesia, Federated states of: Present
EPPO, 2014Guam: Present
EPPO, 2014Indonesia: Present
EPPO, 2014Irian Jaya: Present
Williams & Watson, 1988; EPPO, 2014Kalimantan: Present
Ben-Dov, 1994; EPPO, 2014Sulawesi: Present
IIE, 1995; EPPO, 2014Northern Mariana Islands: Present
Beardsley, 1966Papua New Guinea: Present
Williams, 1982; Cox, 1989; Ben-Dov, 1994; EPPO, 2014Philippines: Present
Cox, 1989; Ben-Dov, 1994; EPPO, 2014Taiwan: Present
Cox, 1989; Ben-Dov, 1994; Le, 1968; EPPO, 2014