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Datasheet

Ceroplastes rusci (fig wax scale)

Summary

  • Last modified
  • 05 May 2015
  • Datasheet Type(s)
  • Pest
  • Natural Enemy
  • Preferred Scientific Name
  • Ceroplastes rusci
  • Preferred Common Name
  • fig wax scale
  • Taxonomic Tree
  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta

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Pictures

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PictureTitleCaptionCopyright
Adults on fig stem.
TitleAdults
CaptionAdults on fig stem.
CopyrightU. Aksoy
Adults on fig stem.
AdultsAdults on fig stem.U. Aksoy
C. rusci located mainly on veins.
TitleC. rusci on fig leaves
CaptionC. rusci located mainly on veins.
CopyrightU. Aksoy
C. rusci located mainly on veins.
C. rusci on fig leavesC. rusci located mainly on veins.U. Aksoy
Fig wax scales on citrus leaf.
TitleWax scales
CaptionFig wax scales on citrus leaf.
CopyrightNOVARTIS Crop Protection AG, Basel Switzerland
Fig wax scales on citrus leaf.
Wax scalesFig wax scales on citrus leaf.NOVARTIS Crop Protection AG, Basel Switzerland

Identity

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Preferred Scientific Name

  • Ceroplastes rusci (Linnaeus, 1758)

Preferred Common Name

  • fig wax scale

Other Scientific Names

  • Calypticus hydatis (Costa) Signoret, 1869
  • Calypticus radiatus Costa, 1829
  • Calypticus testudineus Costa, 1829
  • Ceroplastes denudatus Cockerell, 1893
  • Ceroplastes nerii Newstead, 1897
  • Ceroplastes tenuitectus Green, 1907
  • Chermes caricae (Barnard) Boisduval, 1867
  • Coccus artemisiae Rossi, 1794
  • Coccus caricae Bernard; Fernald, 1903
  • Coccus caricae Fabricius, 1794
  • Coccus hydatis Costa, 1829
  • Coccus rusci Linnaeus, 1758
  • Columnea caricae (Fabricius) Targioni Tozzetti, 1867
  • Columnea testudinata Targioni Tozzetti; Targioni Tozzetti, 1868
  • Columnea testudiniformis Targioni Tozzetti; Targioni Tozzetti, 1866
  • Lecanium artemisiae (Rossi) Signoret, 1872
  • Lecanium radiatum (Costa) Walker, 1852
  • Lecanium rusci (Linnaeus) Walker, 1852
  • Lecanium testudineum (Costa) Walker, 1852

International Common Names

  • English: fig scale
  • Spanish: cappareta de la higuera; cochinilla de la higuera
  • French: ceroplaste du figuier; cochenille du figuier; kermes du figuier

Local Common Names

  • Germany: Feigen-schildlaus; Feigen-wachsschildlaus; Schildlaus, Feigen-; Wachsschildlaus, Feigen-
  • Italy: ceroplaste del fico; cocciniglia del fico

EPPO code

  • CERPRU (Ceroplastes rusci)

Taxonomic Tree

Top of page
  • Domain: Eukaryota
  •     Kingdom: Metazoa
  •         Phylum: Arthropoda
  •             Subphylum: Uniramia
  •                 Class: Insecta
  •                     Order: Hemiptera
  •                         Suborder: Sternorrhyncha
  •                             Superfamily: Coccoidea
  •                                 Family: Coccidae
  •                                     Genus: Ceroplastes
  •                                         Species: Ceroplastes rusci

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

CountryDistributionLast ReportedOriginFirst ReportedInvasiveReferencesNotes

ASIA

AfghanistanPresentBen-Dov, 1993
IndiaRestricted distributionEPPO, 2014
-BiharPresentIIE, 1993; EPPO, 2014
-KarnatakaPresentIIE, 1993; EPPO, 2014
-KeralaPresentIIE, 1993; EPPO, 2014
IndonesiaRestricted distributionEPPO, 2014
-Irian JayaPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014
-JavaPresentNHM, 1979
IranPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
IraqPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
IsraelWidespreadPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
JapanAbsent, invalid recordEPPO, 2014
JordanPresentIIE, 1993; EPPO, 2014
LebanonPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
Saudi ArabiaPresentPellizzari & Camporese, 1994; Ben-Dov, 1993
SyriaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
TurkeyPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014
United Arab EmiratesPresentIIE, 1993; EPPO, 2014
VietnamPresentNga et al., 2006

AFRICA

AlgeriaPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
AngolaPresentBen-Dov, 1993
Cape VerdePresentBen-Dov, 1993; IIE, 1993; EPPO, 2014
EgyptPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
EritreaPresentPellizzari & Camporese, 1994; IIE, 1993
EthiopiaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
GhanaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
KenyaPresentIIE, 1993; EPPO, 2014
LibyaPresentIIE, 1993; EPPO, 2014
MaliPresentEPPO, 2014; Muniappan et al., 2012
MoroccoPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
Sao Tome and PrincipePresentIIE, 1993; EPPO, 2014
SenegalPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
SeychellesPresentBen-Dov, 1993; EPPO, 2014
South AfricaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
SpainPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
-Canary IslandsPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
SudanPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
TanzaniaPresentIIE, 1993; EPPO, 2014
TunisiaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
ZambiaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
ZimbabwePresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014

NORTH AMERICA

USA
-FloridaPresentHamon & Mason, 1997

CENTRAL AMERICA AND CARIBBEAN

Antigua and BarbudaPresentBen-Dov, 1993; IIE, 1993; EPPO, 2014
Dominican RepublicPresentIIE, 1993; EPPO, 2014
GuadeloupePresentEPPO, 2014
Puerto RicoPresentBen-Dov, 1993; EPPO, 2014
Saint Vincent and the GrenadinesPresentSchotman, 1989
United States Virgin IslandsPresentBen-Dov, 1993; EPPO, 2014

SOUTH AMERICA

ArgentinaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
BrazilRestricted distributionPellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014
-Rio Grande do SulPresentIIE, 1993; EPPO, 2014
GuyanaPresentBen-Dov, 1993; IIE, 1993; EPPO, 2014
UruguayPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014

EUROPE

AlbaniaPresentPellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
CroatiaPresentIIE, 1993; EPPO, 2014
CyprusWidespreadBen-Dov, 1993; IIE, 1993; Sisman & Ülgentürk, 2010; EPPO, 2014
FranceRestricted distributionPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
-CorsicaPresentBen-Dov, 1993; IIE, 1993; EPPO, 2014
GibraltarPresentIIE, 1993; EPPO, 2014
GreecePresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
-CretePresentEPPO, 2014
HungaryPresentEPPO, 2014
ItalyPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
-SardiniaPresentEPPO, 2014
-SicilyPresentEPPO, 2014
MaltaPresentIIE, 1993; EPPO, 2014
MontenegroPresentEPPO, 2014
PolandPresent only under cover/indoors<L>abanowski, 2009
PortugalPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
-AzoresPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
-MadeiraPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
SpainPresentPellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
-Balearic IslandsPresentIIE, 1993; EPPO, 2014
Yugoslavia (Serbia and Montenegro)PresentIIE, 1993

OCEANIA

AustraliaAbsent, invalid recordEPPO, 2014

Host plants/Plants Affected

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Plant nameFamilyContext
Actinidia chinensis (Chinese gooseberry)ActinidiaceaeMain
AnnonaAnnonaceaeMain
Annona cherimola (cherimoya)AnnonaceaeMain
Annona muricata (soursop)AnnonaceaeMain
Annona reticulata (bullock's heart)AnnonaceaeMain
Annona squamosa (sugar apple)AnnonaceaeMain
CitrusRutaceaeMain
Citrus aurantium (sour orange)RutaceaeMain
Citrus limon (lemon)RutaceaeMain
Citrus x paradisi (grapefruit)RutaceaeMain
Crataegus (hawthorns)RosaceaeOther
Cydonia (quince)RosaceaeOther
Dimocarpus longan (longan tree)SapindaceaeOther
FicusMoraceaeOther
Ficus carica (fig)MoraceaeMain
Gossypium (cotton)MalvaceaeOther
Hedera helix (ivy)AraliaceaeOther
Ilex aquifolium (holly)AquifoliaceaeOther
Ipomoea batatas (sweet potato)ConvolvulaceaeOther
Juncus (rushes)JuncaceaeOther
Litchi chinensis (lichi)SapindaceaeOther
Mangifera indica (mango)AnacardiaceaeOther
Mespilus germanica (medlar)RosaceaeOther
Morus (mulberrytree)MoraceaeOther
Musa (banana)MusaceaeOther
Myrtus communis (myrtle)MyrtaceaeOther
Nephelium lappaceum (rambutan)SapindaceaeOther
Nerium oleander (oleander)ApocynaceaeOther
Persea americana (avocado)LauraceaeOther
Piper nigrum (black pepper)PiperaceaeOther
PistaciaAnacardiaceaeOther
Pistacia vera (pistachio)AnacardiaceaeOther
Platanus (planes)PlatanaceaeOther
Populus (poplars)SalicaceaeOther
Prunus (stone fruit)RosaceaeOther
Psidium guajava (guava)MyrtaceaeOther
Pyrus communis (European pear)RosaceaeOther
RuscusLiliaceaeOther
Vitis vinifera (grapevine)VitaceaeOther

Growth Stages

Top of pageFlowering stage, Fruiting stage, Seedling stage, Vegetative growing stage

List of Symptoms

Top of page
SignLife StagesType

Fruit

discoloration
external feeding
honeydew or sooty mould

Growing point

discoloration
external feeding

Leaves

external feeding
honeydew or sooty mould

Stems

external feeding
honeydew or sooty mould

Whole plant

external feeding

Natural Enemies

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Natural enemyTypeLife stagesSpecificityReferencesBiological control inBiological control on
Anagyrus kamaliParasiteDiab et al., 2014
Aprostocetus ceroplastaeParasiteAdults/Nymphs
Cheletogenes ornatusPredatorAdults/Nymphs
Coccophagus ceroplastaeParasiteAdults/Nymphs
Coccophagus lycimniaParasiteAdults/Nymphs
Eublemma scitulaPredatorAdults/NymphsItalymandarins
Lewia infectoriaPathogenAdults/Eggs/Nymphs
Metaphycus helvolusParasiteDiab et al., 2014
Pachyneuron muscarumParasiteDiab et al., 2014
Paraceraptrocerus italicusParasiteAdults/Nymphs
Scutellista caeruleaPredatorEggsItalymandarins

References

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abanowski G, 2009. Pests of ornamental plants introduced to Polish glasshouses. (Szkodniki roslin ozdobnych zawlekane do Polskich szklarni.) Progress in Plant Protection, 49(4):1714-1723. http://www.progress.plantprotection.pl/pliki/2009/49-4-35.pdf

Anonymous, 1980. List of pests, plant diseases and weeds for the control of which quarantine measures exist. Zashchita Rastenii, 11:20-21.

Balachowsky AS, 1933. Sur la biologie de 'Ceroplastes floridensis' Comst. et sur la repartition géographique des Ceroplastes dans la region palearctique (Hem. Coccidae). Proceedings 5e Congres International d'Entomologie (Paris 1932), 5:79-87.

Bénassy C, Franco E, 1974. On the ecology of Ceroplastes rusci L. (Homoptera, Lecanoidae) in the Alpes-Maritimes. Annales de Zoologie, Ecologie Animale, 6(1):11-39

Ben-Dov Y, 1970. The wax scales of the genus Ceroplastes Gray (Homoptera: Coccidae) and their parasites in Israel. Israel Journal of Entomology, 5:83-92.

Ben-Dov Y, 1988. The scale insects (Homoptera: Coccoidea) of citrus in Israel: diversity and pest status. Proceedings of the Sixth International Citrus Congress. Tel Aviv, Israel, March 6-11, 1988: 1075-1082.

Ben-Dov Y, 1993. A systematic catalogue of the soft scale insects of the world (Homoptera: Coccoidea: Coccidae) with data on geographical distribution, host plants, biology and economic importance. Gainesville, USA: Sandhill Crane Press, Inc., 536 pp.

Benfatto D, 1982. Studies on the chemical control of pests of citrus: Ceroplastes rusci L. (Hemiptera, Coccidae). Annali dell'Istituto Sperimentale per l'Agrumicoltura, 15/16:121-126

Bodenheimer FS, 1951. Citrus Entomology in the Middle East. The Hague, Netherlands: Dr WJ Junk & S Gravenhage.

Boratynski KL, Williams DJ, 1964. Coccoidea. In: Kloet GS, Hinks WD, eds. Checklist of British Insects. 2nd ed. Handbook for the Identification of British Insects, 11:87-94.

Borchsenius NS, 1957. Subtribe mealybugs and scales (Coccoidea). Soft scale insects Coccidae. Vol. IX. (In Russian). Fauna SSSR. Zoologicheskii Institut Academii Nauk SSSR NS, 66:1-493.

Diab N, Asslan L, Basheer AM, 2014. Parasitoids of the soft scale insects (Homoptera: Coccidae) in fruit orchards at al Qunaetera, Syria. Egyptian Journal of Biological Pest Control, 24(2):341-346. http://www.esbcp.org/index.asp

EPPO, 2014. PQR database. Paris, France: European and Mediterranean Plant Protection Organization. http://www.eppo.int/DATABASES/pqr/pqr.htm

Ezzat YM, Hussein NA, 1969. Redescription and classification of the family Coccidae in UAR. (Homoptera: Coccoidea). Bulletin de la Societe Entomologique d'Egypte, 51:359-426.

Fazeli MJ, Farzneh A, 1993. Biology of Ceroplastes rusci L. on figs in Fars province. Applied Entomology and Phytopathology, 60(1&2):ar 13-17, 3-4

Green EE, 1917. Observations on British Coccidae: With descriptions of new species. Entomologist's Monthly Magazine, 53:201-210, 260-269.

Hamon AB, Mason GJ, 1997. Fig wax scale, Ceroplastes rusci (L.), in Florida (Homoptera: Coccoidea: Coccidae). Entomology Circular (Gainesville), No. 380:2 pp.; 6 ref.

Hodgson CJ, 1994. The scale insect family Coccidae: an identification manual to genera. Wallingford, UK: CAB International, 639 pp.

IIE, 1993. Distribution Maps of Pests, Series A No. 373 (1st revision). Wallingford, UK: CAB International.

Kfir R, Rosen D, 1980. Parasites of soft scales (Homoptera: Coccidae) in Israel: an annotated list. Journal of the Entomological Society of Southern Africa, 43(1):113-128

Muniappan R, Watson GW, Vaughan L, Gilbertson R, Noussourou M, 2012. New records of mealybugs, scale insects, and whiteflies (Hemiptera: Sternorrhyncha) from Mali and Senegal. Journal of Agricultural and Urban Entomology, 28(1):1-7. http://scentsoc.org/Volumes/JAUE/28/28001.pdf

Nga Thi Vu, Eastwood R, Chat Thi Nguyen, Lam Van Pham, 2006. The fig wax scale Ceroplastes rusci (Linnaeus) (Homoptera: Coccidae) in south-east Vietnam: pest status, life history and biocontrol trials with Eublemma amabilis Moore (Lepidoptera: Noctuidae). Entomological Research, 36(4):196-201. http://www.blackwell-synergy.com/loi/enr

Onder EP, Soydanbay M, 1984. Investigations on methods of the chemical control of the wax scale (Ceroplastes rusci L.) damaging fig trees in the pgean Region. Bitki Koruma Bulteni, 24(4):200-212

Onder EP, Ulu O, Zumreoglu A, Saltabas N, 1985. Investigations on methods of chemical control of the fig wax scale (Ceroplastes rusci L.) causing damage on citrus trees in the pgean region. Turkiye Bitki Koruma Dergisi, 9(4):247-253

Pellizzari G, Camporese P, 1994. The Ceroplastes species (Homoptera: Coccoidea) of the Mediterranean Basin with emphasis on C. japonicus Green. Annales de la Societe Entomologique de France, 30(2):175-192

Ragab ME, 1995. Efficiency of Scutellista cyanea Motsch. (Hym., Pteromalidae) and Tetrastichus ceroplastae (Gir.) (Hym., Eulophidae) in population suppression of Ceroplastes rusci L. (Hom., Coccidae). Journal of Applied Entomology, 119(9):627-630; 19 ref.

Scaltriti GPellizzari, Antonucci A, 1982. Notes on some insects injurious to Actinidia crops. (Note su alcuni insetti dannosi a colture di Actinidia.) Informatore Fitopatologico, 32(3):47-48.

Schotman CYL, 1989. Plant pests of quarantine importance to the Caribbean. RLAC-PROVEG, No. 21:80 pp.

Shabana YM, Ragab ME, 1997. Alternaria infectoria, a promising biological control agent for the fig wax scale, Ceroplastes rusci (Homoptera: Coccidae), in Egypt. Biocontrol Science and Technology, 7(4):553-564; 27 ref.

Sisman S, Ülgentürk S, 2010. Scale insects species (Hemiptera: Coccoidea) in the Turkish Republic of Northern Cyprus. Turkish Journal of Zoology, 34(2):219-224. http://journals.tubitak.gov.tr/zoology/

Williams DJ, Watson GW, 1990. The scale insects of the tropical South Pacific region. Part 3: the soft scales (Coccidae) and other families. Wallingford, UK: CAB International, 267 pp.

Distribution Maps

Top of page
Distribution map United Arab Emirates: Present
IIE, 1993; EPPO, 2014United Arab Emirates: Present
IIE, 1993; EPPO, 2014Afghanistan: Present
Ben-Dov, 1993Antigua and Barbuda: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Albania: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Angola: Present
Ben-Dov, 1993Argentina: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Brazil: Restricted distribution
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Brazil
See regional map for distribution within the countryCape Verde: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Cyprus: Widespread
Ben-Dov, 1993; IIE, 1993; Sisman & Ülgentürk, 2010; EPPO, 2014Cyprus: Widespread
Ben-Dov, 1993; IIE, 1993; Sisman & Ülgentürk, 2010; EPPO, 2014Dominican Republic: Present
IIE, 1993; EPPO, 2014Dominican Republic: Present
IIE, 1993; EPPO, 2014Algeria: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Algeria: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Egypt: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Eritrea: Present
Pellizzari & Camporese, 1994; IIE, 1993Spain: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Spain: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Spain
See regional map for distribution within the countrySpain
See regional map for distribution within the countryEthiopia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014France: Restricted distribution
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014France
See regional map for distribution within the countryGhana: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Gibraltar: Present
IIE, 1993; EPPO, 2014Guadeloupe: Present
EPPO, 2014Greece: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Greece: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Greece
See regional map for distribution within the countryGreece
See regional map for distribution within the countryGuyana: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Guyana: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Croatia: Present
IIE, 1993; EPPO, 2014Hungary: Present
EPPO, 2014Indonesia: Restricted distribution
EPPO, 2014Indonesia: Restricted distribution
EPPO, 2014Indonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIndonesia
See regional map for distribution within the countryIsrael: Widespread
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Israel: Widespread
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014India: Restricted distribution
EPPO, 2014India
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIndia
See regional map for distribution within the countryIraq: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Iraq: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Iraq: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Iran: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Iran: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Iran: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Italy: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Italy
See regional map for distribution within the countryItaly
See regional map for distribution within the countryItaly
See regional map for distribution within the countryJordan: Present
IIE, 1993; EPPO, 2014Jordan: Present
IIE, 1993; EPPO, 2014Kenya: Present
IIE, 1993; EPPO, 2014Lebanon: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Lebanon: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Lebanon: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Libya: Present
IIE, 1993; EPPO, 2014Morocco: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Morocco: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Montenegro: Present
EPPO, 2014Mali: Present
EPPO, 2014; Muniappan et al., 2012Malta: Present
IIE, 1993; EPPO, 2014Malta: Present
IIE, 1993; EPPO, 2014Poland: Present only under cover/indoors
<L>abanowski, 2009Puerto Rico: Present
Ben-Dov, 1993; EPPO, 2014Puerto Rico: Present
Ben-Dov, 1993; EPPO, 2014Portugal: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Portugal
See regional map for distribution within the countryPortugal
See regional map for distribution within the countryPortugal
See regional map for distribution within the countrySaudi Arabia: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993Saudi Arabia: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993Seychelles: Present
Ben-Dov, 1993; EPPO, 2014Sudan: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Senegal: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Sao Tome and Principe: Present
IIE, 1993; EPPO, 2014Syria: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Syria: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Syria: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Tunisia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Tunisia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Turkey: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Turkey: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Turkey: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Tanzania: Present
IIE, 1993; EPPO, 2014USA
See regional map for distribution within the countryUSA
See regional map for distribution within the countryUruguay: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Saint Vincent and the Grenadines: Present
Schotman, 1989United States Virgin Islands: Present
Ben-Dov, 1993; EPPO, 2014Vietnam: Present
Nga et al., 2006Yugoslavia (Serbia and Montenegro): Present
IIE, 1993South Africa: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Zambia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Zimbabwe: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
  • = Present, no further details
  • = Evidence of pathogen
  • = Widespread
  • = Last reported
  • = Localised
  • = Presence unconfirmed
  • = Confined and subject to quarantine
  • = See regional map for distribution within the country
  • = Occasional or few reports
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Distribution map (asia) United Arab Emirates: Present
IIE, 1993; EPPO, 2014Afghanistan: Present
Ben-Dov, 1993Indonesia: Restricted distribution
EPPO, 2014Irian Jaya: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Java: PresentIsrael: Widespread
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014India: Restricted distribution
EPPO, 2014Bihar: Present
IIE, 1993; EPPO, 2014Kerala: Present
IIE, 1993; EPPO, 2014Karnataka: Present
IIE, 1993; EPPO, 2014Iraq: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Iran: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Jordan: Present
IIE, 1993; EPPO, 2014Lebanon: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Saudi Arabia: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993Syria: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Turkey: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Vietnam: Present
Nga et al., 2006
Distribution map (europe) Albania: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Cyprus: Widespread
Ben-Dov, 1993; IIE, 1993; Sisman & Ülgentürk, 2010; EPPO, 2014Algeria: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Spain: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Balearic Islands: Present
IIE, 1993; EPPO, 2014France: Restricted distribution
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Corsica: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Gibraltar: Present
IIE, 1993; EPPO, 2014Greece: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Crete: Present
EPPO, 2014Croatia: Present
IIE, 1993; EPPO, 2014Hungary: Present
EPPO, 2014Iraq: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Iran: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Italy: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Sicily: Present
EPPO, 2014Sardinia: Present
EPPO, 2014Lebanon: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Morocco: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Montenegro: Present
EPPO, 2014Malta: Present
IIE, 1993; EPPO, 2014Poland: Present only under cover/indoors
<L>abanowski, 2009Portugal: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Azores: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Madeira: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Syria: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Tunisia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Turkey: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Yugoslavia (Serbia and Montenegro): Present
IIE, 1993
Distribution map (africa) United Arab Emirates: Present
IIE, 1993; EPPO, 2014Angola: Present
Ben-Dov, 1993Cape Verde: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Cyprus: Widespread
Ben-Dov, 1993; IIE, 1993; Sisman & Ülgentürk, 2010; EPPO, 2014Algeria: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Egypt: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Eritrea: Present
Pellizzari & Camporese, 1994; IIE, 1993Spain: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Canary Islands: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Ethiopia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Ghana: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Greece: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Crete: Present
EPPO, 2014Israel: Widespread
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Iraq: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Iran: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Sicily: Present
EPPO, 2014Jordan: Present
IIE, 1993; EPPO, 2014Kenya: Present
IIE, 1993; EPPO, 2014Lebanon: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Libya: Present
IIE, 1993; EPPO, 2014Morocco: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Mali: Present
EPPO, 2014; Muniappan et al., 2012Malta: Present
IIE, 1993; EPPO, 2014Madeira: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014Saudi Arabia: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993Seychelles: Present
Ben-Dov, 1993; EPPO, 2014Sudan: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Senegal: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Sao Tome and Principe: Present
IIE, 1993; EPPO, 2014Syria: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Tunisia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Turkey: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Tanzania: Present
IIE, 1993; EPPO, 2014South Africa: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Zambia: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Zimbabwe: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; IIE, 1993; EPPO, 2014
Distribution map (north america) Dominican Republic: Present
IIE, 1993; EPPO, 2014Puerto Rico: Present
Ben-Dov, 1993; EPPO, 2014Florida: Present
Hamon & Mason, 1997
Distribution map (central america) Antigua and Barbuda: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Dominican Republic: Present
IIE, 1993; EPPO, 2014Guadeloupe: Present
EPPO, 2014Guyana: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Puerto Rico: Present
Ben-Dov, 1993; EPPO, 2014Florida: Present
Hamon & Mason, 1997Saint Vincent and the Grenadines: Present
Schotman, 1989United States Virgin Islands: Present
Ben-Dov, 1993; EPPO, 2014
Distribution map (south america) Argentina: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014Brazil: Restricted distribution
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014Rio Grande do Sul: Present
IIE, 1993; EPPO, 2014Guyana: Present
Ben-Dov, 1993; IIE, 1993; EPPO, 2014Uruguay: Present
Pellizzari & Camporese, 1994; IIE, 1993; EPPO, 2014
Distribution map (pacific) Indonesia: Restricted distribution
EPPO, 2014Irian Jaya: Present
Pellizzari & Camporese, 1994; Ben-Dov, 1993; EPPO, 2014