Biology: Larvae hatch between mid-April and mid-May. Right
after hatching, larvae move to a young shoot and enter through an
internode in the middle of the stem. At the beginning of their development
up to 7 larvae can live together in the same shoot. Larvae of the
twin-spotted wainscot molt five times, mining in 3 to 4 shoots before
pupating in June/July in shoots. After the first change of shoot,
only one larva per shoot can be found. Adults emerge between mid July
and mid August, and live for one to two weeks. Females lay up to 240
eggs in rows under the leaf sheaths of common reed shoots (picture
47).
Description: Larva up to 4.7 cm long;
larval body whitish with several black spots on each segment; head
capsule and parts of the neck shield and anal plate dark-brown/black
(picture 52). Pupa 17 to 24 mm long, reddish-brown, head upwards.
Adults brown winged, with two white spots on the forewings (picture
40).
Feeding pattern/damage: Freshly
hatched larvae enter an internode in the middle of a shoot and at
this entry point sever the shoot tip from the base within 24 hours
(picture 67). Afterwards, the first three instars feed only above
the growing point. Often not only the shoot tips but also the entire
shoots are killed. Such shoots are all smaller than 1 m and can be
recognized by a tiny entrance hole (0.4-0.5 mm wide) on an internode,
the distinct severance of the internode with the entrance hole right
above a node, and a 1-1.3 mm wide exit hole above the growing point.
However, the tip is often broken off above the severance. Feces can
only be found above the growing point. From the first change of shoot
in the third instar onwards, larvae feed also in the upper shoot internodes
which are partly packed with feces. The shoot tips are distinctly
dead (picture 6), but shoots often develop sideshoots on 1-4 nodes
below the damage. Exit holes are always above entrance holes. Holes
made by mature larvae are 2-2.5 mm wide. A plugged entrance hole nearly
2 mm wide and an emergence window (6x3 mm) prepared by the larva indicate
the presence of a pupal chamber (picture 59). Pupae can be found head
upwards in undamaged internodes of lower shoot parts of both damaged
and undamaged shoots (minimum base diameter 5 mm).
Damage caused by the twin-spotted wainscot is very similar to that
of the other three stem-mining wainscots.
Distribution: Europe
References: Bretherton
et al., 1983; Tscharntke,
1990; Häfliger et al.,
2006 |
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Pic. 47: Eggs of Archanara gemini-
puncta
Pic. 52: Mature larva of Archanara
geminipuncta

Pic. 67: Internode severed by first instar larva of A. geminipuncta
Pic. 40: Adult of Archanara gemini-
puncta
Pic. 59: Internode with open emergence window of Archanara geminipuncta
Pic. 6: Shoot tip killed by A. geminipuncta
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